Drassodella septemmaculata ( Strand, 1909 ),

Mbo, Zingisile & Haddad, Charles R., 2019, A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae), Zootaxa 4582 (1), pp. 1-62: 50-53

publication ID

https://doi.org/10.11646/zootaxa.4582.1.1

publication LSID

lsid:zoobank.org:pub:9DC61794-4BD7-4F6D-BB8C-84D9855C8151

persistent identifier

http://treatment.plazi.org/id/B74F87AB-9E50-4015-53B3-CCA462929CA7

treatment provided by

Plazi

scientific name

Drassodella septemmaculata ( Strand, 1909 )
status

 

Drassodella septemmaculata ( Strand, 1909) 

Figs 3View FIGURES 1–4, 12, 16View FIGURES 5–17, 75View FIGURES 60–77, 91View FIGURES 78–92, 167, 168View FIGURES 157–171, 193–196View FIGURES 193–196

Prosthesima  7- maculata Strand, 1909: 554  (Holotype ♀: SOUTH AFRICA: Western Cape: Signal Hill, Cape Town, 33°55'S, 18°24'E, IV.1900, leg. W.F. Purcell, SAM-ENW-X008521—examined).

Drassodella  7- maculata: Tucker, 1923: 314  , fig. 41A, B (♂).

Drassodella septemmaculata: Platnick 1990b: 4  , figs 164–166 (♀ spinnerets).

Diagnosis. Females of D. septemmaculata  are similar to those of D. trilineata  sp. nov., but can be recognised by the shape of the anterior hood, which takes the form of an equilateral triangle, and the heart-shaped posterior half of the median septum ( Figs 75View FIGURES 60–77, 193View FIGURES 193–196), while in D. trilineata  sp. nov. the anterior apex of the anterior hood is broadly rounded and the posterior half of the median septum is almost parallel-sided ( Figs 76View FIGURES 60–77, 197View FIGURES 197–202). Males share with D. vasivulva  the large median apophysis with a retrolaterally-directed basal process, but differ in the shape of the distal end: in D. septemmaculata  it is rounded ( Figs 91View FIGURES 78–92, 195View FIGURES 193–196) and in D. vasivulva  it has an almost perpendicular bend distally ( Figs 92View FIGURES 78–92, 201View FIGURES 197–202).

Description. Female (Signal Hill, NCA 2004/357). Measurements: CL 2.72, CW 2.10, AL 3.56, AW 2.00, TL 6.36 (5.98–6.40), FL 0.38, SL 1.48, SW 1.28, CH 0.10, AME-AME 0.07, AME-ALE 0.02, ALE-ALE 0.27, PME-PME 0.10, PME-PLE 0.13, PLE-PLE 0.48, PERW 0.56, MOQAW 0.21, MOQPW 0.24, MOQL 0.30.

Length of leg segments: I 1.78 + 0.92 + 1.44 + 1.40 + 1.00 = 6.54, II 1.50 + 0.82 + 1.24 + 1.36 + 1.04 = 5.96, III 1.52 + 0.76 + 0.92 + 1.52 + 1.04 = 5.76, IV 2.20 + 1.00 + 1.66 + 2.42 + 1.12 = 8.40.

General appearance as in Fig. 167View FIGURES 157–171. Carapace dark brown, with dense white feathery setae on eye region, lateral sides of cephalic region and towards posterior margin. Clypeus height equal to 1.44 AME diameter; AME smaller than ALE; AME separated by distance slightly less than their diameter, AME separated from ALE by distance equal to 0.25 AME diameter; PME smaller than PLE; PME separated by distance equal to 1.11 their diameter, PME separated from PLE by distance equal to 1.11 PME diameter. Chelicerae dark brown; promargin with three teeth, proximal and median teeth almost touching, median clearly separated from distal tooth; retromargin with two widely spaced subequal teeth. Endites dark brown, pale-yellow at maxillar hair tuft. Labium dark brown. Sternum dark brown, with black mottling, setae more concentrated around margins. Legs dark orange-brown, proximal segments with dense black mottling, faint on metatarsi, absent on tarsi. Leg spination: femora: I do 1 rl 1, II do 2 rl 1, III do 5, IV do 3; patellae: spineless; tibiae: I & II spineless, III pl 2 do 2 rl 2 plv 2 rlv 2 vt 2, IV pl 2 do 2 rl 2 plv 2 rlv 2 vt 2; metatarsi: I pl 1, II plv 1 rlv 1, III pl 4 do 1 rl 4 plv 2 rlv 2 vt 2, IV pl 4 do 2 rl 4 plv 2 rlv 2 vt 2; palpal spination: femora do 3; patellae do 1; tibiae do 3 rl 3; tarsi pl 1 do 3 rl 2. Abdomen black; dorsum with white feathery setae forming three pairs of white markings dorsolaterally, single spot above spinnerets ( Fig. 167View FIGURES 157–171); anterior markings an elongate stripe, second and third pairs of markings spots; oval lateral sclerite above booklungs; venter black, with sparse setae; two paired rows of tiny sclerites, inner rows slightly closer to each other to outer rows. Spinnerets dark brown. Epigyne with shallow anterior hood, forming equilateral triangle, lateral ends terminating abruptly; lateral margins of median septum initially directed mesally, curving in sharp C-shape, posterior two-thirds heart-shaped, converging at epigastric furrow; ST II yellow, connecting ducts and ST I pale yellow-brown; ST II elongate, parallel-sided with rounded head, separated by nearly double their width, nearly twice as large as spherical ST I ( Figs 75View FIGURES 60–77, 193, 194View FIGURES 193–196).

Male (Cederberg Wilderness Area, NCA 2012/961). Measurements: CL 2.52, CW 1.96, AL 2.32, AW 1.36, TL 4.80 (4.76–7.12), FL 0.32, SL 1.32, SW 1.18, CH 0.13, AME-AME 0.07, AME-ALE 0.02, ALE-ALE 0.25, PME-PME 0.11, PME-PLE 0.11, PLE-PLE 0.44, PERW 0.54, MOQAW 0.21, MOQPW 0.23, MOQL 0.30.

Length of leg segments: I 1.78 + 0.98 + 1.32 + 1.28 + 0.84 = 6.20, II 1.70 + 0.84 + 1.24 + 1.30 + 0.98 = 6.06, III 1.56 + 0.72 + 1.10 + 1.44 + 1.12 = 5.94, IV 2.24 + 1.04 + 1.64 + 2.32 + 1.28 = 8.52.

General appearance as in Figs 3View FIGURES 1–4 and 168View FIGURES 157–171. Carapace red-brown, with radiating black mottling; upsilon marking formed by white feathery setae, originating medially from anterior margin, diverging behind fovea towards posterolateral margin. Clypeus height equal to 2.17 AME diameter; AME smaller than ALE; AME separated by distance slightly less than their diameter, AME separated from ALE by distance equal to 0.25 AME diameter; PME smaller than PLE; PME separated by distance equal to 1.33 their diameter, PME separated from PLE by distance equal to 1.33 PME diameter. Chelicerae red-brown anteriorly, reddish posteriorly; promargin with three teeth, proximal and median teeth nearly touching, large space between median and distal teeth; retromargin with two widely separated subequal teeth. Endites brown medially, black along lateral margins, creamy-white around maxillar hair tuft. Labium dark brown medially and posteriorly, light brown anteriorly. Sternum brown. Legs with brown proximal segments, distal segments orange. Leg spination: femora: I do 2, II do 2, III do 5, IV do 3; patellae: spineless; tibiae: I & II spineless, III pl 2 do 2 rl 2 plv 2 rlv 2 vt 2, IV pl 2 do 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I & II spineless, III pl 2 rl 2 plv 2 rlv 2 vt 2, IV pl 4 do 2 rl 4 plv 2 rlv 2 vt 2; palpal spination: femora do 1; patellae do 1 rl 1; tibiae do 3 rl 3 rlv 1; tarsi pl 1 do 1 rl 1 plv 1. Abdomen black dorsally, with small orange-brown scutum extending quarter abdomen length ( Fig. 168View FIGURES 157–171); dorsum with white feathery setae forming three pairs of white markings dorsolaterally, single spot above spinnerets; anterior markings an elongate stripe, contiguous with second spotted markings, third pair isolated spots; elongate lateral sclerite above booklungs; venter creamy-yellow; one pair of tiny sclerites medially, running from booklungs to midpoint of abdomen; small post-epigastric sclerites present, ante-epigastric sclerites absent. Spinnerets cream. Palp deep orange-brown; RTA long, entirely obscured in ventral view by retrolateral heel of cymbium; in retrolateral view triangular, elongate, with narrow base, gradually tapering to rounded tip; cymbium narrow, 2.11 times longer than wide, narrowed to rounded point distally; embolus originating prolaterally, narrow, with undulating curves and sharp tip, directed retrodistally; median apophysis massive, generally Y-shaped, with prominent rounded retrolateral lobe, longer prolaterally-directed finger-shaped tip partly obscuring embolus ( Figs 91View FIGURES 78–92, 195, 196View FIGURES 193–196).

Additional material examined. SOUTH AFRICA:  Western Cape Province: Bergvliet, Diep River , Cape Flats, 34°03'S, 18°27'E, VIIGoogleMaps  .1905, leg. W.F. Purcell, 1♀ ( SAMC ENW-B003343)  ; Cape Town, Constantia, 34°02'S, 18°23'E, 16. IIGoogleMaps  .1992, leg. B. Heydenrych (pitfall, fynbos near summit), 1♂ ( MRAC 173621View Materials)  ; Cape Town, Gardens, 33°56'S, 18°24'E, VIIGoogleMaps  .1898, leg. E.A. Morris, 1♀ ( SAMC ENW-X004330)  ; Cape Town, Muizenberg, 34°06'S, 18°27'E, 1989, leg. R. Legg (pitfalls), 11♂ ( MRAC 169869View Materials)GoogleMaps  , 3♂ 7♀ ( MRAC 169870View Materials)GoogleMaps  , 11♀ ( MRAC 169871View Materials)  ; Same locality, 5.V.1991, leg. R. Legg (dunes to the north), 8♀ ( MRAC 173713View Materials)  ; Same data as previous but 19.V.1991, 11♀ ( MRAC 173722View Materials); Same data as previous but 2. VI  .1991, 3♀ ( MRAC 173738View Materials); Same data as previous but 16. VI  .1991, 5♀ ( MRAC 173768View Materials)  ; Cape Town, Newlands, 33°58'S, 18°29'E, V.1899, leg. W.F. Purcell, 1♀ ( SAMC ENW-X006000)GoogleMaps  ; Cape Town, Newlands Forest , Paradijs Stream, 33°57'S, 18°27'E, 7. IIIGoogleMaps  .1992, leg. N. Larsen (leaf litter), 1♀ ( SAMC ENW-C002494)  ; Cape Town, Signal Hill, 33°56'S, 18°24'E, IV–VGoogleMaps  . 1899, leg. W.F. Purcell, 4♀ ( SAMC ENW-X005978); Same locality, VII  .1901, leg. W.F. Purcell, 1♀ ( SAMC ENW- X008974)  ; Same locality, V.1902, leg. W.F. Purcell, 1♀ ( SAMC ENW-X 012082)  ; Cape Town, Table Mountain , Maclears Beacon, 33°58'S, 18°25'E, VIGoogleMaps  .1904, leg. R.M. Lightfoot, 1♀ ( SAMC ENW-X013593)  ; Cape Town, Table Mountain National Park , Signal Hill, 33°54'S, 18°22'E, 13. VIIGoogleMaps  .1978, leg. N. Larsen (hand, under stones in grass tussocks), 2♀ ( NCA 2004 /357)  ; Cape Town, Table Mountain , ravine below Platteklip, 33°57'S, 18°25'E, IVGoogleMaps  .1900, leg. W.F. Purcell, 1♀ ( SAMC ENW-X008563)  ; Cape Town, Table Mountain woods, 28. VIII  .1915, leg. R.W. Tucker, 1♀ ( SAMC ENW-B001340); Cederberg Wilderness Area, Wupperthal, Crystal Pools , 32°27.540'S, 19°08.334'E, 1285 m a.s.l.GoogleMaps  , X.2009, leg. E. Nortje & S. Kritzinger-Klopper (pitfall traps), 1♂ ( NCA 2012 /961); Cederberg Wilderness Area , Wupperthal, Crystal Pools, 32°19.932'S, 19°08.460'E, 1135 m a.s.l., IIIGoogleMaps  .2009, leg. E. Nortje & S. Kritzinger-Klopper (pitfall traps), 1 subadult ♀ ( NCA 2012 /3776); Cederberg Wilderness Area , Sneeukop, 32°21.216'S, 19°09.174'E, 1615 m a.s.l., IIIGoogleMaps  .2009, leg. E. Nortje & S. Kritzinger-Klopper (pitfall traps), 1♂ ( NCA 2011 /2712); Cederberg Wilderness Area , Sneeukop, 32°21.324'S, 19°09.150'E, 1605 m a.s.l., IIIGoogleMaps  .2010, leg. E. Nortje & S. Kritzinger-Klopper (pitfall traps), 1♂ ( NCA 2011 /2736); Same data as previous but IIIGoogleMaps  .2005, 1♂ ( NCA 2012 /2204); Cederberg Wilderness Area , Sneeukop, 32°21.330'S, 19°09.150'E, 1845 m a.s.l., IIIGoogleMaps  .2008, leg. E. Nortje & S. Kritzinger-Klopper (pitfall traps), 1♂ ( NCA 2012 /3537); Same data as previous but IIIGoogleMaps  .2007, 1♂ ( NCA 2012 /3774); Cederberg Wilderness Area , Sneeukop, 32°21.330'S, 19°09.924'E, 1880 m a.s.l., IIIGoogleMaps  .2007, leg. E. Nortje & S. Kritzinger-Klopper (pitfall traps), 1♂ ( NCA 2012 /3775); False Bay , St James, 34°07'S, 18°27'E, IVGoogleMaps  .1899, leg. W.F. Purcell, 1♀ ( SAMC ENW-X005968); Same locality, IV  .1901, leg. Dr Purcell, 3♀ ( SAMC ENW-X009195)  ; Fisherhaven, 34°21.430'S, 19°07.557'E, 12.I.2008, leg. C. Haddad (leaf litter), 1♂ ( NCA 2008 / 452); Great Winterhoek Mountains , 33°08'S, 19°10'E, 4000–5000 ft a.s.l., 4–14. IVGoogleMaps  .1916, leg. R.W. Tucker, 1♀ ( SAMC ENW-B002550); Hout Bay , 34°02'S, 18°21'E, IIIGoogleMaps  .1898, W.F. Purcell, 1♂ ( SAMC ENW-X004344); Side of Kalk Bay and St James Mountain , 34°08'S, 18°27'E, 27. IIGoogleMaps  .1902, leg. W.F. Purcell, 2♂ 3 imm. ( SAMC ENW- X 012311); Tulbagh, Piquetberg Road Station , 33°22'S, 19°07'E, 9. VIIIGoogleMaps  .1898, leg. R.M. Lightfoot, 1♀ ( SAMC ENW-X003998)  . Northern Cape Province: Oorlogskloof Nature Reserve , Nieuwoudtville, 31°31'S, 19°26'E, 29. IVGoogleMaps  .2006, leg. J. Leroy (renosterveld area), 1♀ ( NCA 2007 /1305)  .

Distribution. Known from various localities in the Western and Northern Cape Provinces, South Africa ( Fig. 203View FIGURE 203).

SAMC

Iziko Museums of Cape Town

VI

Mykotektet, National Veterinary Institute

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Gallieniellidae

Genus

Drassodella

Loc

Drassodella septemmaculata ( Strand, 1909 )

Mbo, Zingisile & Haddad, Charles R. 2019
2019
Loc

Drassodella

Tucker, R. W. E. 1923: 314
Loc

Drassodella septemmaculata: Platnick 1990b : 4

Platnick, N. I. 1990: 4