Arboricaria sociabilis Kulczynski , 1897

Mikhailov, Kirill G., 2016, On the spider genus Arboricaria with the description of a new species (Araneae, Gnaphosidae), ZooKeys 558, pp. 153-169: 162-164

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Arboricaria sociabilis Kulczynski , 1897


Taxon classification Animalia Araneae Gnaphosidae

Arboricaria sociabilis Kulczynski, 1897   Figs 12-16, 17-20

Micaria sociabilis   Kulczyński in Chyzer & Kulczyński, 1897: 254 & 255 (key), 258-259, Tab.X., figs 21 (♀) 25 a–b (♂).

Micaria canestrinii   Roewer, 1951: 447 (replacement name for Micaria aurata   Canestrini, 1868, praeocc.).

Micaria sociabilis   : Wunderlich 1980: 291-292, figs 36 a–b (♂, doubtful, see note in the text below, incorrect drawings), 60a (♀).

Arboricaria koeni   Bosmans in Bosmans & Blick, 2000: 465, figs 32-35 (♂♀), syn. n.

Micaria sociabilis   : Pfliegler 2014: 145 (record), figs 3e, 4c (♂♀).

Micaria sociabilis   : Sentenská et al. 2015: figs 4, 6 (♂♀).

Not Micaria sociabilis   : Wunderlich 1980: figs 60 b–c (♀, = Arboricaria brignolii   , see below).


1 ♂ ( IRSNB, holotype of Arboricaria koeni   ), Greece, Kreta, Chania , [in bark], 22.V.1994 (leg. Koen van Keer)   ; 1 ♂ ( IRSNB, paratype of Arboricaria koeni   , left palp missing), Kreta, Chania, 22-5-1994   ; 1 ♂ ( ZMMU; left palp and leg I only), Russia, Rostov-on-Don , 47°13'33"N, 39°41'59"E, window of living flat, 8.06.1978 (leg. et det. A.V. Ponomarev) GoogleMaps   .

Taxonomic remarks.

Originally, the male was matched with the female with some doubts ( Chyzer and Kulczyński 1897), because they were taken from different, but not extremely distant localities of the former Austro-Hungarian Empire. Syntypes (1 ♂, 1 ♀, “Ungarn” = “Hungary”) are listed by Wunderlich (1980), but he only redescribed the female. Comparing the epigynes of Arboricaria sociabilis   and Arboricaria koeni   shows no essential difference between them; therefore, these names are to be synonymized. The position of the copulatory openings is a little variable; in the type of Arboricaria sociabilis   , they are closer to the middle part of the epigynal groove, in the Arboricaria koeni   type and the Arboricaria sociabilis   material as depicted by Pfliegler (2014) closer to the posterior one-third.

A male syntype of Micaria sociabilis   from Mukachevo is currently kept in the Zoological Museum in Warsaw, Poland, but both palps are missing (W. Wawer, pers. comm.). The tibial apophysis as redrawn by Wunderlich (1980: Fig. 36a, see also Fig. 14) from the original description ( Chyzer and Kulczyński 1897: tab. X, fig. 25b, see also Fig. 12) is certainly incorrect. No deep bifurcation is visible in the original figure. Miller (1971) in his key to Czechoslovak spiders, pointed out: "Tibial apophysis apically [sic! - KM] forked with 2 teeth, lower tooth narrower and more pointed; it is laterally slightly bent, ventrally rounded outside and bent forward and it has the same width" (translated from Czech by A. Šestaková). Miller’s specimen of Micaria sociabilis   was never depicted and is currently missing among the other Micaria   samples kept in the National Museum in Prague, Czech Republic ( Kůrka 1994).

The picture of the Micaria sociabilis   male palp as presented by Pfliegler (2014) certainly indicates the identity of this species with Arboricaria koeni   (male types examined, see above in Material) and additionally confirms the synonymy of these names. To be ex act, not enough details are visible in the publication, but a correct shape of the embolus is shown in the original photograph kindly sent to me by the author (cf. Figs 15 and 16 with traced embolus).

Characteristically, a male specimen from Rostov-on-Don was initially identified by A.V. Ponomarev as Micaria sociabilis   , only later re-labeled as Arboricaria koeni   .


Ukraine: Transcarpathia: Mukachevo (= Munkácz in Chyzer & Kulczyński [1897]). NE-Hungary (two other localities from the original description; Debrecen [ Pfliegler 2014]); Spain, continental France, together with Corsica, Italy, Croatia, Macedonia, continental Greece, together with Crete, Bulgaria, Romania, Czech Republic, Slovakia ( Helsdingen 2014, for Micaria sociabilis   and Micaria koeni   ). Russia: Rostov Area ( Ponomarev and Tsvetkov 2006a, as Arboricaria cyrnea   ; Ponomarev 2008; Ponomarev and Dvadnenko 2013), Krasnodar Province: Kushchevskaya ( Ponomarev and Tsvetkov 2006b, as Arboricaria brignolii   ; Ponomarev 2008). Azerbaijan (Caucasus Major: Huseynov, Alieva, 2010, as Arboricaria koeni   , Apsheron Peninsula: Huseynov 2002, as Arboricaria koeni   ), all for Micaria   (or Arboricaria   ) koeni   . The records of Arboricaria cyrnea   from the Rostov Area, as well as those of Arboricaria brignolii   from the Rostov Area and Krasnodar Province belong to Arboricaria sociabilis   (A.V. Ponomarev, pers. comm., as Arboricaria koeni   ).


See Sentenská and Pekár (2013), Sentenská et al. (2015).


Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique
















Arboricaria sociabilis Kulczynski , 1897

Mikhailov, Kirill G. 2016

Arboricaria koeni

Bosmans 2000

Arboricaria brignolii

Bosmans 2000