Soyedina washingtoni (Claassen, 1923)

Grubbs, Scott A. & Baumann, Richard W., 2019, Soyedina Ricker, 1952 (Plecoptera: Nemouridae) in the eastern Nearctic: review of species concepts, proposed morphology-based species groups, and description of a new species from North Carolina, Zootaxa 4658 (2), pp. 223-250 : 231

Soyedina washingtoni Group

http://lsid.speciesfile.org/urn:lsid: Plecoptera .speciesfile.org:TaxonName:50713 6

Three species are proposed for this group: S. carolinensis , S. merritti , and S. washingtoni . In ventral aspect the epiproct dorsal sclerite is open distally, revealing a sclerotized inner member which is flared apically, directed slightly to the left, and the flange lip is approximately the same size radially ( Figs. 26–28 , 36–38 , 44–46 ). The paired ventral sclerites are asymmetric with right member distinctly longer than left ( Figs. 25–26 , 35–36 , 43–44 ). The paired dorsal sclerites are also asymmetric ( Figs. 29–30 , 39–40 , 47–48 ). The outer paraproct lobes are variable between species. This is mainly an Appalachian-distributed species group ( Fig. 80 ).

Figures

FIGURE 80. Distribution maps of eastern Nearctic Soyedina based on material examined during this study plus valid literature data. 80, S. carolinensis, S. merritti, and S. washingtoni.

FIGURES 25–30. Soyedina carolinensis. 25: USA, Virginia, seep into Lewis Fork; 26, 28: USA, West Virginia, Marsh Fork of Buckhannon River; 27, 30: USA, North Carolina, Rattlesnake Springs; 29: USA, North Carolina, Robin Branch; 25, epiproct, ventral view, 170X; 26, epiproct, ventral view, 370X; 27, epiproct apex, ventral view, 650X; 28, epiproct apex, dorsal view, 1200X; 29, epiproct, lateral view, 285X; 30, epiproct, caudal view, 230X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member.

FIGURES 35–42. Soyedina merritti. 35–38, 40–41: USA, Pennsylvania, seep into Powdermill Run; 39, 42: USA, Pennsylvania, seep into Indian Creek. 35, epiproct, ventral view, 150X; 36, epiproct, ventral view, 250X; 37, epiproct apex, ventral view, 650X; 38, epiproct apex, dorsal view, 550X; 39, epiproct, lateral view, 270X; 40, epiproct, caudal view, 220X; 41, outer paraproct lobe, lateral view, 180X; 42, outer paraproct lobe, lateral view, 250X. ds = dorsal sclerite, vs = ventral sclerite, im = inner member.

FIGURES 43–50. Soyedina washingtoni. 43–46, 49: USA, Maine, Thorndike; 47: USA, Maryland, spring into Elk Lick Run; 48: USA, Pennsylvania, Fourmile Creek; 50: USA, Maryland, Mill Run. 43, epiproct, ventral view, 105X; 43, epiproct, ventral view, 247X; 45, epiproct apex, ventral view, 637X; 46, epiproct apex, dorsal view, 488X; 47, epiproct, lateral view, 370X; 48, epiproct, caudal view, 250X; 49, outer paraproct lobe, lateral view, 166X; 50, outer paraproct lobe, lateral view, 300X; ds = dorsal sclerite, vs = ventral sclerite, im = inner member.