Upeneus heterospinus Uiblein & Pavlov, 2019

Uiblein, Franz, Williams, Jeffrey T., Bailly, Nicolas, Hoang, Tuan A. & Rajan, P. Thomas, 2024, Four new goatfishes (Upeneus, Mullidae, Mulliformes) from the Asian Indo-Pacific with a list of valid goatfish species and remarks on goatfish diversity, Cybium 48 (2), pp. 135-160 : 160

Upeneus heterospinus Uiblein & Pavlov, 2019

Types. Holotype: VNMN-I 2015 , 91 mm SL, South-central Vietnam, W Pacific, South China Sea, Khanh Hoa province, Nha Trang, N of Hon Tre, 12°13’21”N, 109°18’30”E, obtained from hookah divers; paratypes: South-central Vietnam, W Pacific, South China Sea: Phu Yen province, Vung Ro Bay, 12°52.11’N, 109°24.66’E, obtained from hookah divers: VNMN-I 2016 , 70 mm SL; VNMN-I 2017, 101 mm SL; VNMN-I 2018 , 96 mm SL; Khanh Hoa province , Nha Trang Bay : N of Hon Tre, 12°13’21”N, 109°18’30”E, obtained from hookah divers: VNMN-I 2020 , 97 mm SL; VNMN-I 2021, 112 mm SL; VNMN-I 2022, 106 mm SL; VNMN-I 2023, 113 mm SL; VNMN-I 2024, 105 mm SL; VNMN-I 2025, 109 mm SL; VNMN-I 2026, 100 mm SL; VNMN-I 2027, 112 mm SL; VNMN-I 2028, 104 mm SL; VNMN-I 2029, 104 mm SL; VNMN-I 2030, 103 mm SL; VNMN-I 2031 , 96 mm SL; VNMN-I 2032, 103 mm SL; S of Hon Tre, obtained from hookah divers: HIFIRE F 58110, 120 mm SL; HIFIRE F 58115, 109 mm SL; VNMN-I 2033, 123 mm SL; Cua Be : CSIRO H 8427-01 , 90 mm SL, landing site, 12°12.09’N, 109°12.12’E; HIFIRE F 58203, 105 mm SL, fish market, 12°12.10’N, 109°12.05’E; Nha Trang city, fish market: CSIRO H 8428-01 , 82 mm SL; CSIRO H 8428-02 , 82 mm SL; HIFIRE F 58178, 152 mm SL; NHMO J 7217, 114 mm SL; NHMO J 7218 ,

103 mm SL; Nha Trang, no further locality details: MNHN 1965- 272, 90 mm SL; ZMUC P49483, 84 mm SL.

Non-types. N Vietnam, W Pacific, South China Sea, Qu ảng Ninh province , Ha Long Bay , Van Don landing site, 21°03.61’N, 107°25.81’E: HIFIRE F 58382 , 97 mm SL; NHMO J 7219, 104 mm SL; VNMN-I 2034, 109 mm SL; VNMN-I 2035, 113 mm SL; VNMN-I 2036, 104 mm SL; VNMN-I 2037, 108 mm SL; VNMN-I 2038, 108 mm SL; Ha Long: VNMN-I 2039 , 83 mm SL; VNMN-I 2040 , 83 mm SL; VNMN-I 2041, 122 mm SL; VNMN-I 2042 , 93 mm SL; VNMN-I 2043, 108 mm SL; VNMN-I 2044 , 94 mm SL; VNMN-I 2045, 117 mm SL; VNMN-I 2046 , 87 mm SL; VNMN-I 2047 , 72 mm SL; VNMN-I 2048 , 82 mm SL; VNMN-I 2049 , 80 mm SL; VNMN-I 2050 , 77 mm SL; VNMN-I 2051 , 81 mm SL; South-central Vietnam, Khanh Hoa province, Nha Trang Bay : HIFIRE F 58231, 123 mm SL, N of Hon Tre Island, 12°14’18”N 109°15’54”E, obtained from hookah divers; Gulf of Thailand, S Vietnam, Phu Quoc, An Thoi market, 10°0.90’N, 104°0.86’E: HIFIRE F 58287 , 93 mm SL; NHMO J 7220 , 88 mm SL; VNMN-I 2054 , 83 mm SL; VNMN-I 2055, 114 mm SL; Gulf of Thailand, Thailand, Ko-Sichang Island : CAS 17679 , 40 (of 141): 66-94 mm SL, 13°8’N, 100°51’E, trawl, 18-24 m depth ; Singapore: NHMO J 2134 , 2 , 78-81 mm SL; Philippine Sea, Central Philippines, Panay Island : UPVMI 155 , 96 mm SL ; Indonesia, Bali Sea, E Java: CSIRO H 7360-04 , 1 (of 3), 66 mm SL, Banguwangi , 08°09’S 114°23’E; Lombok (border of W Pacific to Indian Ocean): CSIRO H 8409-02 , 65 mm SL, and CSIRO H 7364-02 , 127 mm SL, Tanjung Luar, 8°45’S, 116°35’E; BPBM 29896 , 96 mm SL .