Afroestricus, Scarbrough Table Of Contents, 2005

Scarbrough Table Of Contents, Aubrey G., 2005, Afroestricus, a new Afrotropical Ommatine (Diptera: Asilidae) genus with twenty species and two species groups, Zootaxa 1041 (1), pp. 1-76 : 8-11

Afroestricus , gen. n. ( Figs. 1–196 , Tables 1, 2)

Type species: Ommatius macroscelis Bezzi 1906: 294 , by present designation.

Discription: Small to medium sized flies (6.3–20.6 mm) with brown to black body, wings entirely tomentose, and distinctive terminalia ( Figs. 5–192 ), especially the roof­like epiphallus which conceals distiphallus dorsally and tubular spermatheca. Head: Ventral 2/3 of face ( Fig. 1 ) moderately gibbose. Palpus and proboscis yellow setose. Antenna, frons, and ocellar tubercle brown to black setose. Frons with wide stripe of long, multiple setae laterally; sides slightly converging dorsally. Ocellar tubercle with two long setae, each about as long as basal three antennomeres combined.

Thorax: Scutellum usually with two black marginal bristles and prominent preapical marginal groove. Stout anepimeral bristle present. Wing ( Figs. 3a–3b ): Entirely microtrichose, usually brownish­yellow, sometimes brown or yellow. Legs: Fore femur ( Fig. 2 ) with preapical, anteroventral bristle. Hind femur with two ventral rows of bristles, each extending from base to apex; posteroventral row of bristles usually arched medially onto posterior surface. Tarsi partially yellow, at least a few yellow bristles present. Apical segments of abdomen of females usually with sparse, unusually stout, bristles. Terminalia (5–192). Male: Cerci unusually long and narrow, three or more times as long as wide. Epandrium notched dorsally or apically, with a short flat flange along internal surface apically. Aedeagal sheath strongly produced [=epiphallus] anteriorly, concealing tubular distiphallus dorsally and usually apex laterally; distiphallus usually horizontal, apex somewhat flattened laterally, often pointed and recurved; base of aedeagus produced anteriorly, projecting between gonocoxites and beyond hypandrium, usually wide and rounded. Female: Spermatheca tubular, recurved except in A. minutus , usually slightly flattened and wider apically, sides gradually wider apically; spermathecal duct unusually narrow, including valves, each connected to large bulbus bursa basally ( Fig. 24 ) and spermatheca preapically. Sternum 8 with minute spurlike ridge apically. Genital fork mostly membranous, arms sclerotized dorsally.

Etymology. Afroestricus is a combination of the prefix Afro for the Afrotropical region where the species occurs and the Latin estricis for eating in excess, referring to the voracious feeding habits of asilids. The gender is feminine.

Remarks. Afroestricus is distinguished from Ommatius by the moderately gibbose face; frons with wide stripe of long setae, not arranged in single file; ocellar setae subequal to or as long as three antennal segments combined ( Fig. 1 ); fore femur with anterior, preapical bristle ( Fig. 2 ); the flat, rooflike epiphallus which conceals the tubular distiphallus dorsally; the notched epandrium and the tubular, recurved, slightly flattened spermatheca.

Figures

FIGURES 1–2. Character states of Afroestricus Scarbrough. 1. Head of Afroestricus chiastonureus (Speiser) in lateral view. 2. Apex of fore femur of Afroestricus chiastonureus (Speiser) showing a preapical anteroventral bristle. Abbreviations: Ocellar seta=Oc; Preapical anteroventral bristle= Pab.

FIGURES 3–4. Character states of the wing and postocular bristles of the subgenus Afroestricus Scarbrough. 3a–b. Wing venation of the macroscelis and the minutus species groups. 4a–b. Short, erect and long, thin, postocular bristles bristles. Abbrevations: First radial cell=r1; fourth radial cell=r4; discal cell=d; radial­medial crossvein=r­m; first medial cell=m1; third medial cell=m3.

FIGURES 5–14. Afroestricus abyssinia, sp. n. Male terminalia: 5–7. Lateral, dorsal, and ventral views. 8. Gonostylus. 9–10. Aedeagus, dorsal and lateral views. Female terminalia: 11–12. Dorsal and ventral views. 13. Spermatheca. 14. Genital fork. Abbreviations: Aedeagus=Ae; cercus=Ce; epandrium=Ep; epandrial flange=Ef; gonocoxite=Gc; lateral and median corner of gonocoxite= L, M; gonostylus=Gs; median corner of gonocoxite; epiphallus=Eph; hypandrium=Hp; tergum 8=T8; tergum 9=T9; sternum 8=St8.

FIGURES 15–24. Afroestricus chiastoneurus (Speiser). Male terminalia: 15–17. Lateral, dorsal, and ventral views. 18. Gonostylus. 19–20. Aedeagus, dorsal and lateral views. Female terminalia: 21–22. Dorsal and ventral views. 23. Spermatheca. 24. Genital fork with bulbus bursa and three spermathecal ducts.

FIGURES 25–34. Afroestricus eminentis, sp. n. Male terminalia: 25–27. Lateral, dorsal, and ventral views. 28. Gonostylus. 29–30. Aedeagus, dorsal and lateral views. Female terminalia: 31–32. Dorsal and ventral views. 33. Spermathecae. 34. Genital fork.

FIGURES 35–44. Afroestricus fulaui, sp. n. Male terminalia: 35–37. Lateral, dorsal, and ventral views. 38. Gonostylus. 39–40. Aedeagus, dorsal and lateral views. Female terminalia: 41–42. Dorsal and ventral views. 43. Spermathecae. 44. Genital fork.

FIGURES 45–54. Afroestricus hamulus, sp. n. Male terminalia: 45–47. Lateral, dorsal, and ventral views. 48. Gonostylus. 49–50. Aedeagus, dorsal and lateral views. Female terminalia: 51–52. Dorsal and ventral views. 53. Spermatheca. 54. Genital fork.

FIGURES 55–64. Afroestricus insectaris, sp. n. Male terminalia: 55–57. Lateral, dorsal, and ventral views. 58. Gonostylus. 59–60. Aedeagus, dorsal and lateral views. Female terminalia: 61–62. Dorsal and ventral views. 63. Spermatheca. 64. Genital fork.

FIGURES 65–70. Afroestricus kimerus, sp. n. Male terminalia: 65–67. Lateral, dorsal, and ventral views. 68. Gonostylus. 69–70. Aedeagus, dorsal and lateral views.

FIGURES 71–80. Afroestricus macroscelis (Bezzi). Male terminalia: 71–73. Lateral, dorsal, and ventral views. 74. Gonostylus. 75–76. Aedeagus, dorsal and lateral views. Female terminalia: 77–78. Dorsal and ventral views. 79. Spermatheca. 80. Genital fork.

FIGURES 81–90. Afroestricus minutus (Bromley). Male terminalia: 81–83. Lateral, dorsal, and ventral views. 84. Gonostylus. 85–86. Aedeagus, dorsal and lateral views. Female terminalia: 87–88. Dorsal and ventral views. 89. Spermatheca. 90. Genital fork.

FIGURES 91–100. Afroestricus morani, sp. n. Male terminalia: 91–93. Lateral, dorsal, and ventral views. 94. Gonostylus. 95–96. Aedeagus, dorsal and lateral views. Female terminalia: 97–98. Dorsal and ventral views. 99. Spermatheca. 100. Genital fork.

FIGURES 101–110. Afroestricus persuasus (Oldroyd). Male terminalia: 101–103. Lateral, dorsal, and ventral views. 104. Gonostylus. 105–106. Aedeagus, dorsal and lateral views. Female terminalia: 107–108. Dorsal and ventral views. 109. Spermatheca. 110. Genital fork.

FIGURES 111–120. Afroestricus sadaukii, sp. n. Male terminalia: 111–113. Lateral, dorsal, and ventral views. 114. Gonostylus. 115–116. Aedeagus, dorsal and lateral views. Female terminalia: 117–118. Dorsal and ventral views. 119. Spermatheca. 120. Genital fork.

FIGURES 121–130. Afroestricus sankofa, sp. n. Male terminalia: 121–123. Lateral, dorsal, and ventral views. 124. Gonostylus. 125–126. Aedeagus, dorsal and lateral views. Female terminalia: 127–128. Dorsal and ventral views. 129. Spermatheca. 130. Genital fork.

FIGURES 131–140. Afroestricus variabilis (Engel). Male terminalia: 131–133. Lateral, dorsal, and ventral views. 134. Gonostylus. 135–136. Aedeagus, dorsal and lateral views. Female terminalia: 137–138. Dorsal and ventral views. 139. Spermatheca. 140. Genital fork.

FIGURES 141–150. Afroestricus varipes (Curran). Male terminalia: 141–143. Lateral, dorsal, and ventral views. 144. Gonostylus. 145–146. Aedeagus, dorsal and lateral views. Female terminalia: 147–148. Dorsal and ventral views. 149. Spermatheca. 150. Genital fork.

FIGURES 151–160. Afroestricus velatus, sp. n. Male terminalia: 151–153. Lateral, dorsal, and ventral views. 154. Gonostylus. 155–156. Aedeagus, dorsal and lateral views. Female terminalia: 157–158. Dorsal and ventral views. 159. Spermatheca. 160. Genital fork.

FIGURES 161–166. Afroestricus verutus, sp. n. Male terminalia: 161–163. Lateral, dorsal, and ventral views. 164. Gonostylus. 165–166. Aedeagus, dorsal and lateral views.

FIGURES 167–176. Afroestricus victus, sp. n. Male terminalia: 167–169. Lateral, dorsal, and ventral views. 170. Gonostylus. 171–172. Aedeagus, dorsal and lateral views. Female terminalia: 173–174. Dorsal and ventral views. 175. Spermatheca. 176. Genital fork.

FIGURES 177–186. Afroestricus vittatus (Curran). Male terminalia: 177–179. Lateral, dorsal, and ventral views. 180. Gonostylus. 181–182. Aedeagus, dorsal and lateral views. Female terminalia: 183–184. Dorsal and ventral views. 185. Spermatheca. 186. Genital fork.

FIGURES 187–192. Afroestricus vorax, sp. n. Male terminalia: 187–189. Lateral, dorsal, and ventral views. 190. Gonostylus. 191–192. Aedeagus, dorsal and lateral views.