Parhomonyx Ohaus, 1915

Moore, Matthew R., Jameson, Mary L., Garner, Beulah H., Audibert, Cedric, Smith, Andrew B. T. & Seidel, Matthias, 2017, Synopsis of the pelidnotine scarabs (Coleoptera, Scarabaeidae, Rutelinae, Rutelini) and annotated catalog of the species and subspecies, ZooKeys 666, pp. 1-349 : 1

Parhomonyx Ohaus, 1915 Figs 1C , 51

Type species.

Homonyx fuscoaeneus Ohaus, 1905.

Species.

1 species; length 17-22 mm.

The monotypic genus Parhomonyx is endemic to northern Argentina. Ohaus (1905) described P. fuscoaeneus in the genus Homonyx and in conjunction with another unusual pelidnotine, E. oryctoides (originally Homonyx orcytoides ). According to Ohaus (1915b), the genera Homonyx and Parhomonyx were closely related, and differences in the mandibular form (apex with two teeth in Homonyx versus apex rounded in Parhomonyx ) provide character support for both genera (=lineages). Species in the pelidnotine genera Parhomonyx , Homonyx , and Pseudogeniates are distributed primarily in the southern half of South America. Ohaus (1915b) considered Parhomonyx to be an "intermediate stage" that "led Homonyx to Pseudogeniates " ( Ohaus 1915b: 258), and that characters such as coloration, clypeus, mouthparts, elytra, metatibia, and antennae indicated a progression of forms ( Jameson and Ocampo 2012). Additional research should examine sister-group relationships of the taxon.

Parhomonyx fuscoaeneus is castaneous-bronze in color and is diagnostic for its rounded mandibular apex with apical tooth (shared with Pseudogeniates ; bidentate in Homonyx ); metatibial apex with many spinules (biemarginate in Homonyx ); elytral apex with a fringe of setae (shared with Pseudogeniates ; hidden in Homonyx ); protibia lacking weak, basal notch; all claws simple; fifth meso- and metatarsomeres with one or two internal teeth (lacking in Pseudogeniates , shared with Homonyx ); lateral set of setae on apical edge of 3rd metatarsomere of equal length and width (versus unequal length and width in Pseudogeniates ); pronotum with bead complete apically, laterally, and basally; prosternal process short; mesosternal peg lacking (shared with Homonyx and Pseudogeniates ); elytra striate (shared with Pseudogeniates and Homonyx ); and body form elongate and parallel-sided (shared with Homonyx and Pseudogeniates ). Larvae are not described. Label data indicate that specimens are collected at blacklight.

Figures

Figure 1. Clypeal shape varying from rounded, parabolic, trapezoidal, subquadrate (A-D), and emarginate (E-F). Lateral edge of mandibles with no reflexed teeth (lacking teeth in A, but the mandible is reflexed rather than flattened in A, two reflexed teeth (B, E, F), or one reflexed tooth (C )). Apical bead of pronotum varying from obsolete (A) to complete medially (B, C, D, F). Disc of frons with V-shaped depression (E) or frons planar, smooth, lacking V-shaped depression (A-D, F) A Chrysina beyeri Skinner B Epichalcoplethis velutipes velutipes (Arrow) C Parhomonyx fuscoaeneus (Ohaus) D Chipita mexicana (Ohaus) E Sorocha sp. F Homonyx elongatus (Blanchard).

Figure 51. Homonyx fuscoaeneus Ohaus (valid name Parhomonyx fuscoaeneus [Ohaus]) male type (see " Type specimens and lectotype designation " in Methods) from ZMHB. A Dorsal habitus B Lateral habitus C Specimen labels, mouthparts, and male genitalia D Male genitalia, lateral view E Male parameres, dorsal view.