Anotylus sculpturatus ( Gravenhorst, 1806 )

Wang, Lin-Fei, Zhou, Hong-Zhang & Lü, Liang, 2017, Revision of the Anotylus sculpturatus group (Coleoptera: Staphylinidae: Oxytelinae) with descriptions of seven new species from China, Zootaxa 4351 (1), pp. 1-79 : 57-58

Anotylus sculpturatus ( Gravenhorst, 1806)

(Figs. 16)

Gravenhorst, 1806: 187 ( Oxytelus ; variety of carinatus; Type locality: Not cited); Thomson, 1859: 45 (type species of Anotylus ); Mulsant & Rey, 1878: 514 ( Oxytelus ; subgenus Anotylus ; larval characters; France); Lohse, 1964: 85 ( Oxytelus ; subgenus Anotylus ; central Europe); Herman, 1970: 419 ( Anotylus ); Herman, 2001: 1391 ( Anotylus ; catalog); Smetana, 2004: 515 ( Anotylus ; Palaearctic catalog; distribution); Schülke & Smetana, 2015: 773 ( Anotylus ; Palaearctic catalog; distribution).

Syn.: Oxytelus flavipes Lacordaire, 1835: 464 (nec Stephens, 1834, preoccupied; Type locality: Paris); Erichson, 1839: 592 ( Oxytelus ; synonym of sculpturatus ); Fowler, 1888: 380 ( Oxytelus ; synonym of sculpturatus ).

Oxytelus terrestris Heer, 1839: 205 (nec Lacordaire, 1835, preoccupied; Type locality: Matt, auch in den Alpen, Frugmatt); Kraatz, 1857: 855 ( Oxytelus ; synonym of sculptus); Ganglbauer, 1895: 641 ( Oxytelus ; synonym of sculpturatus ).

Oxytelus montivagus Heer, 1841: 574 (Type locality: Nufenen, 5000’); Fauvel, 1872: 171 [= 1873: 12] ( Oxytelus ; synonym of sulpturatus).

Oxytelus havelkai Dvořák, 1954: 38 (subgenus Anotylus ; aberration of sculpturatus ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Boheme centr. Zbraslav); Herman, 2001: 1393 ( Anotylus ; Nomen nudum; catalog).

Oxytelus belgicus Dvořák, 1954: 38 (subgenus Anotylus ; variety of sculpturatus ; Type locality: Belgique: Mouscron); Herman, 2001: 1393 ( Anotylus ; synonym of sculpturatus ; catalog).

Oxytelus coussementi Dvořák, 1954: 40 (subgenus Anotylus ; aberration of sculpturatus variety belgicus ; [Note: Nomen nudum (Articles 1.3.4, 45.5)]; cited from Belgique: Mouscron); Herman, 2001: 1393 ( Anotylus ; Nomen nudum; catalog).

Material examined. EUROPE: 13 exs., Cro. Bernh Portoré 97// Chicago NHMus M. Bernhauer Collection// FMNHINS 2821258–1270 Field Museum ( FMNH); 1 male, same data, FMNHINS 2822156 Field Museum ( FMNH); 1 male, Cro. Bernh Portoré 97// sculpturatus Gr. det. Bernhauer// Chicago NHMus M. Bernhauer Collection// FMNHINS 2821255 Field Museum ( FMNH); 1 male, same data, FMNHINS 2822157 Field Museum ( FMNH); 9 exs., Chichester, Engl. // from Rev. C. E. Tottenham// FMNHINS 2822291–99 Field Museum. ( FMNH); 4 exs., SWITZERLAND: Rüttenen (Solothurn)// 1–8. VI. 1976 B. and H. Malkin// FMNHINS 2822213– 16 Field Museum ( FMNH); 1 female, same data, FMNHINS 2822211 Field Museum ( FMNH); 1 male, HOMF TAUERN KOETSCHACHTAL // Chicago NHMus M. Bernhauer Collection// FMNHINS 2821287 Field Museum ( FMNH); 1 male, Hohe Tauern Nassfeld // Chicago NHMus M. Bernhauer Collection// FMNHINS 2821286 Field Museum ( FMNH); 1 male, Salzburg Lofer// sculpturatus Gravh. // Chicago NHMus M. Bernhauer Collection// FMNHINS 2821288 Field Museum ( FMNH); 1 male, GASTEINERTAL BERNHAUER// ANGERTAL BOO M. VI. 1930 ERLENLAUB// Chicago NHMus M. Bernhauer Collection// FMNHINS 2821293 Field Museum ( FMNH).

Redescription. Body black, fairly shiny; elytra brownish, with base relatively dark; legs brown. Forebody pubescent, densely punctate and moderately striate. Body length [average] male, 4.31 mm; female, 3.68 mm.

Male: Head (Fig. 16-1C, 16-2A) slightly broader than pronotum. Disc convex, with large and round punctures and dense longitudinal striae laterally; mid-longitudinal sulcus short and deep, with deep impression on each side before base. Supra-antennal ridge with margin incrassate. Temples dilated, approximately two times as long as eyes. Clypeus nearly square, about 1/3 of total head length, slightly convex and coriaceous, with a few small punctures; anterior margin slightly rounded. Epistomal suture with lateral portions straight and parallel, running backward to level of anterior margin of eyes.

Occipital suture curved forward in middle. Antenna filiform, article IV globular, V–X short and transverse, with ultimate article stilliform. Neck with smooth area carrying a few punctures, same size as on head.

Pronotum (Fig. 16-1D, 16-2A) transverse, broadest at near anterior 1/3, with relatively dense punctures and striae, even denser laterally; disc slightly convex, mid-longitudinal sulcus straight and deep, changed narrow and deep to base, with two shallow, narrow paramedial sulci, and bounded by obtuse longitudinal carina, with shallow depression close to lateral margins. Anterior margin slightly emarginate in middle. Anterior angles obtuse and slightly produced. Lateral and posterior margins running continuously, nearly curve-formed, posterior angles not obvious. Scutellum with crest-shaped impression, sharply pointed with median lobe. Elytra densely striate and punctate, not pubescent; lateral margin curved and posterior margins truncate and slightly slant. Abdomen coriaceous, covered with punctures and pubescence. Sternite VII (Fig. 16-1G, 16-2C) with posterior margin almost straight and bearing long setae laterally, in middle with a pair of small tubercles, beyond posterior margin; Sternite VIII (Fig. 16-1H, 16-2D) with posterior margin deeply emariginate and carrying long setae laterally.

Aedeagus (Fig. 16-1J–M, 16-3A–C) with median lobe oval, inflated at base and shortly narrowing to apex; apical orifice large and ventral orifice crescentic; internal sac of median lobe with complicated sclerite. Parameres (Fig. 16-1N, O, 16-3D) closely situated to each other and twining around median lobe, with base half flattened and enlarged, apex half narrow and elongate, with thick seta and two rounded processes at middle of ventral edge of each paramere.

Female. Head (Fig. 16-1E, 16-2B) smaller than male, narrower than pronotum. Temples shorter, as long as eyes. Pronotum (Fig. 16-1F, 16-2B) with sculptures coarser. Sternite VII (Fig. 16-2E) with posterior margin weekly emarginate. Sternite VIII (Fig. 16-1I, 16-2F) with posterior margin shallowly broadly bi-emarginate rectangular in middle and protruded. Spermatheca (Fig. 16-1P, 16-3E) with base bulb-like, with apical part clavate and slightly curved.

Distribution. Mongolia; Russia; Kazakhstan; Iran; Cyprus; Turkey; Europe; Algeria.