Tomarus maimon Erichson, 1847

López-García, Margarita M., Giraldo-Mendoza, Alfredo E. & Deloya, Cuauhtémoc, 2019, Taxonomic synopsis of the genus Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae) in Peru, Zootaxa 4604 (3), pp. 428-440 : 432-433

Tomarus maimon Erichson, 1847

( Figs. 13–15 )

Tomarus maimon Erichson, 1847: 96 Heteronychus fossator Burmeister, 1847: 101 . Designed as a subspecies by Dupuis (2016) and then synonymized by López- García & Deloya (2018).

Diagnosis. Tomarus maimon is characterized by the antennal club subequal in length to antennomeres 2–7; clypeus narrowed towards apex (base 2.5 times wider than apex); frontoclypeal tubercles conical; pronotal tubercle acute and prominent; pronotal fovea rounded (1/ 3x as wide as interocular distance); protibial tridentate, without an additional basal denticle; apex of metatibia with 16–20 spinules; parameres with a small, acute tooth each side, apical half strongly contracted and slender, apices expanded ( Fig. 14 ).

Locality records ( Fig. 15 ). 107 males and 94 females from CEUC, MNHUB, FSCA, MEKRB, SENASA, and USNM . 11 specimens from CMNC. Ancash (1): Santa, Moro. Arequipa (1): Camaná , Quilca. Cajamarca

(1): Baños del Inca. Ica (36): Chavín, Cerro Lindo; Chincha; Ica; Paracas, Santa Cruz; Topará; Villacurí. Junín (12): Chanchamayo; La Merced; Pichanaqui; Satipo, Mazamari. La Libertad (15): Ascope, Chócope; Libertad; Pacasmayo; Trujillo; Trujillo, Hacienda Casa Grande; Virú. Lambayeque (2): Chiclayo, Lambayeque. Huánuco (7): Aucayacu, Tingo María. Lima (87): Canta, Santa Rosa de Quives; Cañete; Huaura, Lachay; Huaura, Sayán/Andahuasi; Huaral, Huando; Huarochirí, Matucana; Huarochirí, San Pedro de Casta; Huarochirí, Santa Eulalia; Lima, La Molina ; Lurín; Quebrada Verde. Loreto (11): Alto Amazonas, Yurimaguas. Yanamono. Moquegua (1): Ilo, Río Osmore. Piura (11): Chira; Morropón, Chulucanas; Paita; Piura. Tambogrande/Progreso Bajo. Tacna (6): Jorge Basadre, Locumba; Tacna. San Martín (1): Tocache. Imprecise data (20): Peru .

Temporal distribution. January (10), February (8), March (1), April (8), May (20), June (7), July (23), August (7), September (16), October (28), November (23), December (23). No data (38).

Distribution. Ecuador, Peru, Chile, Brazil, and the Guianas (Endr̂di 1985).

Natural history. Ayquipa & Cueva (1979) cited T. maimon (as Bothynus maimon ) as a pest of sugar cane that affects the roots and stumps. It also attacks the roots of the sweet potato, known as camote in Peru ( Daza & Rincón 1993).

Figures

FIGURES 13–15. Tomarus maimon: 13, habitus; 14, parameres in frontal view; 15, locality records in Peru. Habitus photograph by Yony Callohuari.