Calityini Reitter, 1922, 1758

Kolibac, Jiri, 2013, Trogossitidae: A review of the beetle family, with a catalogue and keys, ZooKeys 366, pp. 1-194 : 8

Tribe Calityini Reitter, 1922

Calityini Reitter, E. 1922: 66.

Type genus.

Calitys Thomson, 1859

Bouchard, P. et al. 2011: 57. Crowson, R. A. 1970: 13 (referred as Calitinae subfam.nov.). Ślipiński, S. A. 1992: 442 ( Calitinae ). Lawrence, J. F. & Newton, A. F., Jr. 1995: 869 ( Calitinae ). Kolibáč, J. 2006: 117 ( Calityni Winkler, 1922; sic!) (diagnosis, new status). Kolibáč, J. 2007a: 364. Kolibáč, J. & Leschen, R. A. B. 2010: 242

Remarks.

The position of the single genus Calitys within the trogossitid system has changed many times over the past century or so. It has been classified within either Peltinae or Trogossitinae (compare, for example, Barron 1971 and Crowson 1964a vs. Crowson 1970), then treated as a separate subfamily ( Crowson 1970 in Trogossitidae s.str. without Peltidae , Ślipiński 1992 in Trogossitidae s.lat.). Reitter’s early idea (1876) that it might be classified within the former Leperini or Leperinae (i.e. Gymnochilini herein) is also interesting and worthy of review. Calitys belongs among the primitive fungivorous groups and has several features shared with Peltinae , for example: robust mandibles with mola, flat body, wide pronotum, weakly asymmetrical antennal club, and absence of elytral interlocking mechanism. However, it also has bizarre sculptures on dorsal surface of body, wax scales, and tufts of rigid setae that together differentiate the genus from all other trogossitids. Its basal position in the trogossitine branch is based chiefly on procoxal cavities perfectly externally closed, presence of paragular sclerites in larval cranium and concave larval tergite IX. Wax scales and tufts of setae on head, antennae, elytra, and pronotum make it resemble Gymnochilini . Nonetheless, it remains possible to imagine Calitys also as a derived member of Peltinae or even Lophocaterinae .