Diplonevra goliatha, Corona, Emily M. & Brown, Brian V., 2005

Corona, Emily M. & Brown, Brian V., 2005, The Central American species of Diplonevra Lioy (Diptera: Phoridae), Zootaxa 1050, pp. 21-38 : 29-30

Diplonevra goliatha new species

Figs. 3 , 16 , 22

Recognition. See the closely related D. ereba , from which this species differs by the lightcolored palpus, forelegs, and halter, as well as the thinner, fewer setae at the base of the inner face of the hind femur ( Fig. 3 ).

Description. Male. Body length 4.40–4.90 mm. Frons black to brown, densely punctate. Mean frontal width 0.51 head width. Flagellomere 1 brown, slightly spherical. Arista brown. Palpus orange, setose with many small, fine setulae along margin laterally; apex with several long, thick pointed setae that range from 0.10–0.20 mm. Proboscis yellow, ventrally setose, elongate, narrow. Scutum black­brown, yellow mediolaterally, brown anterolaterally, with 2 brown anteromedial stripes; scutellum black with 2 pairs of large setae. Pleuron dark brown, tomentose, shiny. Forelegs yellow. Apices of mid­ and hind femora with darkening on both anterior and posterior faces; basal two­thirds yellow. Hind tibia black with 3–4 anterodorsal setae. Large medial and small lateral region of posterior face of hind trochanter with dense, extremely fine setae ( Fig. 3 ). Ventrobasal region of posterior face of hind femur with several thin setae present, lacking process. Wing, brown, especially near wing tip, vein R2+3 present, fork large ( Fig. 16 ). Costal length 0.48–0.49 wing length. Costal setae 0.10–0.15 mm. Mean costal sector ratio 5.1:1.3:1.0; range 5.0–5.2:1.2–1.5:1.0. Base of radial sector with 2 small setae. Halter yellow. Abdominal tergites black­brown. Venter of abdomen dark gray. Left side of epandrium dark brown, tomentose; posterior margin setose, with 1 long and several shorter setae. E pandrium with protruding, transparent lobe. Left side of hypandrium dark brown, tomentose except shiny posterodorsally. Right side of epandrium ( Fig. 22 ) brown, tomentose posteriorly, dark brown dorsally; posterior margin setose, with 1 extremely thick seta ventrally. Cercus yellow, setose. Stalk of cercus yellow, setose except anteroventral one­quarter bare.

Female. Body length 6.40–6.70 mm. Frons black, densely punctate. Swelling with medial indentation above supra­antennal setae present on frons. Mean frontal width 0.51 head width. Flagellomere 1 brown, slightly spherical. Arista brown. Palpus dorsoventrally flattened, orange, elongated extending beyond flagellomere 1. Lateral margin of palpus with small, fine setulae; apex of palpus with several long, thick pointed setae (0.18–0.28 mm). Clypeus elongated, black­brown. Proboscis elongate, geniculate. Labrum glossy, light brown. Labellae glossy, orange. Scutum black­brown except light brown mediolaterally, anterolaterally, 2 anteromedial strips; scutellum black with 2 pairs of large setae. Pleuron black­brown, tomentose, shiny except dorsum of anepisternum black, dorsal two­thirds of meron dark brown, anterior margin of katepisternum light brown. Forelegs yellow except femur dark brown dorsal two­thirds. Mid­ and hind femora with darkening on both anterior and posterior faces; basal half yellow. Hind tibiae black with 3–4 anterodorsal setae. Wing vein R2+3 present, fork large. Costal length 0.49–0.52 wing length. Costal setae 0.15–0.20 mm. Mean costal sector ratio 5.8:1.5:1.0; range 5.6–6.0:1.4–1.6:1.0. Wing membrane noticeably darkened, especially anteriorly. Halter yellow. Abdominal tergites black and present only on segments 1–4. Tergite 1 broad, short. Tergite 2 covers entire segment. Tergite 3 extends full length of segment, but only fourfifths of width. Tergite 4 extends full length of segment, but only three­sevenths of width. Venter of abdomen dark gray.

Geographical distribution. Honduras

Derivation of specific epithet. Derived from the name of the Philistine giant, referring to the large size of this species.

Holotype. ɗ, HONDURAS: Cortes: Parque Nacional Cusuco, 5 km N. Buenos Aires, 15.48°N, 88.22°W, 30. vi. 1995, R. Cave. [ LACM ENT 053932] ( LACM).

Paratypes. HONDURAS: Cortes: Parque Nacional Cusuco, 5 km N. Buenos Aires, 15.48°N, 88.22°W, 1ɗ, 15. ii.1995, 1 Ψ, 30. iii.1995, 1 ɗ, 30. vi.1995, 1 Ψ, 15.vii.1995, R. Cordero, R. Cave, Malaise trap, 1600 m ( LACM, MZLU).

Figures

FIGURES 1 – 6. Diplonevra spp. Basal region of posterior face of hind tibia and trochanter (scanning electron micrographs). Fig. 1. D. ereba new species. Fig. 2. D. gnoma new species. Fig. 3. D. goliatha new species. Fig. 4. D. hypermeka new species. Figs. 5 – 6. D. setigera (Malloch), Zurquí de Moravia, Costa Rica. Abbreviations: dms – dorsomedial setulae of trochanter; ls – lateral setulae of trochanter.

FIGURES 14 – 19. Diplonevra spp. Left wings.

FIGURES 20 – 25. Diplonevra spp. Male terminalia, right lateral.