Reddyanus Vachon, 1972

Kovařík, František, Lowe, Graeme, Ranawana, Kithsiri B., Hoferek, David & Š, V. A., 2016, Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n, Euscorpius 220, pp. 1-133 : 53-56

publication ID	https://doi.org/ 10.18590/euscorpius.2016.vol2016.iss220.1
publication LSID	lsid:zoobank.org:pub:DD0DF45D-F63A-4AA2-8EFF-03CF99E297EF
DOI	https://doi.org/10.5281/zenodo.7124533
persistent identifier	https://treatment.plazi.org/id/B07187DF-A821-FFA9-FED7-F9DB9F33FAC6
treatment provided by	Felipe
scientific name	Reddyanus Vachon, 1972
status

Treatment

Genus Reddyanus Vachon, 1972 View in CoL stat. n.

( Figs. 14 View Figures 12–15 , 199 View Figures 193–200 , 201–220 View Figures 201–208 View Figures 209–224 , 225–236 View Figures 225–240 , 245–250, 254–402 View Figures 241–259 View Figures 260–263 View Figures 264–269 View Figures 270–275 View Figures 276–282 View Figures 283–286 View Figures 287–290 View Figures 295–300 View Figures 301–302 View Figures 303–306 View Figures 307–312 View Figures 313–318 View Figures 319–326 View Figures 327–330 View Figures 331–333 View Figures 334–337 View Figures 338–348 View Figures 349–354 View Figures 355–357 View Figures 358–361 View Figures 362–365 View Figures 366–371 View Figures 372–375 View Figures 376–377 View Figures 378–381 View Figures 382–386 View Figures 387–392 View Figures 393–400 View Figures 401–402 , 409–420 View Figures 403–429 , 551–554 View Figures 547–554 , 561–570 View Figures 555–570 , Tables 3–5 View Table 3 View Table 4 View Table 5 )

Isometrus (Reddyanus) : Vachon, 1972: 176–177, figs. 14, 16, 18, 20; Vachon, 1982: 90–101, 108–110, figs. 47–67, 88–91; Fet & Lowe, 2000: 150–154; Kovařík, 2003: 5–14, figs. 1–8; Kovařík & Ojanguren, 2013: 184–193, 347–354, 357–360, figs. 1265–1367, 1384–1409.

Isometrus View in CoL ( Raddyanus [sic]): Tikader & Bastawade, 1983: 254–311 (in part), figs. 727–770, 841–895.

TYPE SPECIES. Isometrus acanthurus Pocock, 1899 .

DIAGNOSIS. Medium sized buthids, adults 19–75 mm. Sternum type 1, triangular in shape. Pedipalps orthobothriotaxic, type Aβ. Chelal trichobothrium db located between et and est. Three to five pairs of lateral eyes. Tibial spurs absent on all legs. Pedipalp movable finger with six rows of granules, several accessory granules and external and internal accessory granules. Pedipalp fixed finger with seven rows of granules and six external and seven internal accessory granules. Cheliceral fixed finger with a single ventral denticle. Third and fourth legs with tibial spurs. Tibia and tarsomeres of legs I–III with setae not arranged into bristle combs on dorsal surfaces. Tarsomere II of leg IV with two sparse rows of <20 spiniform setae on ventral surface. Mesosoma dorsally with one median carina. Telson with subaculear tooth pointed or more often rounded. Males of most species have longer metasomal segments and often also a wider pedipalp chela manus than females.

COMMENTS. Vachon (1972: 177) described the subgenus Reddyanus and distinguished it from the nominotypic subgenus Isometrus Ehrenberg, 1828 by the position of chelal fixed finger trichobothria (trichobothrium db located between dt and et in Isometrus , between et and est in Reddyanus ). Vachon also cited the different positions of the e 2 trichobothria on the femur (see figs 15, 16 in Vachon, 1972: 176). Subsequently, Vachon (1982) added another character for distinguishing the two subgenera: Isometrus has pedipalp fixed fingers with six rows of granules and six external and internal accessory granules ( Figs. 252–253 View Figures 241–259 and figs. 64, 65 in Vachon, 1982: 100), and Reddyanus has pedipalp fixed fingers with seven rows of granules and six external and seven internal granules ( Figs. 254–259 View Figures 241–259 and figs. 66 and 67 in Vachon, 1982: 100). Here we propose two additional characters for distinguishing these taxa: (i) ventral aspect of tarsomere II of leg IV with two dense rows of> 30 longer, more filamentous setae in Isometrus ( Figs. 196– 197 View Figures 193–200 ), vs. two sparse rows of <20 shorter, thicker, more spiniform setae in Reddyanus ( Figs. 199 View Figures 193–200 , 205–208 View Figures 201–208 ); (ii) hemispermatophore with a relatively short, broad trunk, a short, broad, uncoiled flagellum, and a smaller, blunt basal lobe in Isometrus ( Figs. 146–149 View Figures 146–149 ), vs. a narrow trunk, a long, thin, coiled flagellum and a much enlarged, pointed basal lobe in Reddyanus ( Figs. 283– 286 View Figures 283–286 ). The form of the hemispermatophore of Isometrus appears more similar to that of Lychas ( Figs. 183–186 View Figures 183–186 ), than to Reddyanus . Differences in trichobothriotaxy, pedipalp finger dentition, tarsal setation and hemispermatophore structure provide characters that are useful for higher level classification of buthid scorpions ( Kovařík, 2009; Lamoral, 1979; Sissom, 1990; Stahnke, 1972; Stockwell, 198; Vachon, 1952). Taken together, these differences support our taxonomic decision to elevate Reddyanus to a genus distinct from Isometrus .