Ochodaeus, Dejean, 1821

Paulsen, M. J., 2019, Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae), Insecta Mundi 684, pp. 1-14 : 11

Key to species of Ochodaeus in Madagascar

1. Pronotum everywhere strongly, densely granulate with elevated tubercles ( Fig. 13 )........ 2

— Pronotum either entirely smooth and punctate or with weak granulation confined to disc ( Fig. 14 )........................................................................ 4

2(1). Elytral tubercles on intervals sparse; body length less than 4 mm .... O. pygmaeus Paulian — Elytral tubercles dense; body length greater than 4 mm ............................... 3

3(2). Median thickening of clypeal bead variable, lateral thickening always evident ( Fig. 15 ); body usually> 9.0 mm....................................... O. isoanalensis Paulian

— Median thickening of clypeal bead simply convex, bead thin laterally ( Fig. 16 ); body length usually 4.5–8.0 mm................................................... O. miliaris Klug

4(1). Pronotal surface simply punctate............. O. modopunctatus Paulsen , new species — Pronotal surface weakly granulate on disc.......................................... 5

5(4). Clypeal bead evenly curved ( Fig. 17 ). Mentum deeply, entirely furrowed ( Fig. 5 ). Males usually with 2 pollices on protibia.................. O. polypollicatus Paulsen , new species

— Clypeal bead wavy ( Fig. 18 ). Mentum with furrow not reaching past middle ( Fig. 6 ). Males with a single pollex on protibia...................... O. meandrus Paulsen , new species

Figures

Figures 13–18. Key characters of Madagascar Ochodaeus species. 13) Granulate pronotum of O. isoanalensis Paulian. 14) Punctate pronotum of O. modopunctatus Paulsen, new species. 15) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 16) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin. 17) Head of O. polypollicatus Paulsen, new species, showing smoothly curved clypeal margin. 18) Head of O. meandrus Paulsen, new species, showing wavy clypeal margin.

Figures 4–12. Additional characters of Madagascar Ochodaeus species. 4–6) Mentum, shading indicates foveae. 7–9) Male genitalia, inset with arrow illustrating shape of sclerotized patch on internal sac between parameres. 10–12) Right stridulatory peg, dorsal view. 4, 7, 10) O. modopunctatus Paulsen, new species. 5, 8, 11) O. polypollicatus Paulsen, new species. 6, 9, 12) O. meandrus Paulsen, new species.