Platycerus

Huang, Hao & Chen, Chang-Chin, 2009, Notes on the morphology, taxonomy, and natural history of the genus Platycerus Geoffroy from China, with the description of a new species (Coleoptera: Scarabaeoidea: Lucanidae), Zootaxa 2087, pp. 1-36 : 4-6

Checklist of Platycerus of the world

Platycerus Geoffroy, 1762

= Systenocerus Weise, 1883

= Systenus Sharp & Muir, 1912

Type species: Scarabaeus caraboides Linnaeus, 1758

North American species 1:

1. Platycerus depressus LeConte, 1850 —northeastern USA & Canada = Platycerus piceus Kirby, 1837 (unavailable name) 2

2. Platycerus marginalis Casey, 1897 —western North America = Platycerus cribripennis Van Dyke, 1928 (synonym)

3. Platycerus oregonensis Westwood, 1844 —western North America = Platycerus coerulescens LeConte, 1869 (synonym) = Platycerus chalybaeus Casey, 1897 (synonym)

4. Platycerus virescens (Fabricius, 1775) (Lucanus) —eastern & central USA, Canada

= Lucanus quercus Weber, 1801 (synonym)

= Lucanus securidens Say, 1823 (synonym)

= Platycerus quercus angustus Casey, 1909 (synonym) = Platycerus quercus iowanus Casey, 1909 (synonym) = Platycerus peregrinus Casey, 1909 (synonym)

European and western Asian species 3:

5. Platycerus caprea ( De Geer, 1774) (Lucanus) —Europe

6. Platycerus caraboides (Linnaeus, 1758) (Scarabaeus) — Europe, Syria, Iran, Morocco, Algeria, Tunisia = Platycerus azureus Dalla Torre, 1879 (synonym)

= Platycerus caerulosus Didier & Seguy, 1953 (possible synonym) 4 = Carabus coerulescens Uddman, 1753 (synonym)

= Platycerus chalybaeus MacLeay, 1819 (synonym) 5 = Platycerus chlorizans Dalla Torre, 1879 (synonym) 5 = Platycerus coeruleus Dalla Torre, 1879 (synonym) 5 = Platycerus cribratus Mulsant & Rey, 1863 (synonym) = Platycerus diabolus Dalla Torre, 1879 (synonym) 6 = Platycerus nigripes MacLeay, 1819 (synonym) 6

= Platycerus pseudocaprea Paulus, 1970 (synonym) 7 = Lucanus rufipes Herbst, 1790 (synonym)

= Platycerus trivialis Dalla Torre, 1879 (synonym)

= Platycerus virescens Mulsant, 1842 (synonym)

= Platycerus viridiaeneus Mulsant, 1842 (synonym)

= Platycerus viridicollis Depoli, 1917 (synonym)

= Platycerus viridis Dalla Torre, 1879 (synonym)

7. Platycerus primigenius Weise, 1960 — Turkey, Armenia, Azerbaijan, Iran

8. Platycerus senguni Schweiger, 1966 — Turkey

9. Platycerus spinifer Schaufuss, 1862 — France, Portugal, Spain

10. Platycerus caucasicus Parry, 1864 —Caucasus, Turkey, Georgia, Armenia, Azerbaijan 11. Platycerus delagrangei Fairmaire, 1892 — Turkey, Syria

12. Platycerus perplexus Gusakov, 2003 — Azerbaijan, NE Georgia, Russia 13. Platycerus vicinus Gusakov, 2003 — Georgia, N Armenia

Japanese species:

14. Platycerus acuticollis acuticollis Kurosawa, 1969 — Japan

15. Platycerus acuticollis takakuwai Fujita, 1987 — Japan 8 Platycerus acuticollis akitai Fujita, 1987 — Japan 8 Platycerus acuticollis namedai Fujita, 1987 — Japan 8

16. Platycerus delicatulus delicatulus Lewis, 1883 — Japan Platycerus delicatulus unzendakensis Fujita & Ichikawa, 1982 — Japan

17. Platycerus kawadai Fujita & Ichikawa, 1982 — Japan

18. Platycerus sugitai Okuda & Fujita, 1987 — Japan

19. Platycerus urushiyamai Imura, 2007 — Japan

20. Platycerus akitaorum Imura, 2007 — Japan

21. Platycerus sue Imura, 2007 — Japan

Chinese species ( Fig. 158 ):

22. Platycerus hongwonpyoi hongwonpyoi Imura & Choe, 1989 — South Korea = Platycerus hongwonpyoi merkli Imura & Choe, 1989 (synonym?)— North Korea Platycerus hongwonpyoi mongolicus Imura & Bartolozzi, 2006 —Inner Mongolia (Ao-han-qi) Platycerus hongwonpyoi qinlingensis Imura, 1993 —Shaanxi (Qinling) = Platycerus hongwonpyoi dabashanensis Okuda, 1997 (synonym?)—Chongqing (Dabashan) = Platycerus hongwonpyoi funiuensis Imura, 2006 (synonym?)—Henan (Funiushan) = Platycerus hongwonpyoi shennongjianus Imura, 2008 (synonym?)—Hubei (Shennongjia) Platycerus hongwonpyoi tianmushanus Imura & Wan, 2006 —Zhejiang (Tianmushan)

23. Platycerus tabanai tabanai Okuda & Tanikado, 1994 —Shaanxi (Qinling) = Platycerus tabanai taibaishanensis Okuda, 2003 (synonym?)—Shaanxi (Qinling) Platycerus tabanai baotianmanus Imura, 2006 —Henan (Funiushan)

24. Platycerus rugosus Okuda, 1997 —Chongqing (Dabashan), NE Sichuan (Micangshan)

25. Platycerus yingqii Huang & Chen, 2009 —Shaanxi (Qinling)

26. Platycerus dundai Imura & Bartolozzi, 1994 —W Sichuan (Luding)

= Platycerus benesi Imura & Bartolozzi, 1994 (synonym) 9

Platycerus dundai miyatakei Tanikado & Tabana, 1997 —S Sichuan (Jinyang) 10

27. Platycerus cupreimicans Imura, 2006:121 —Yunnan

28. Platycerus feminatus Tanikado & Tabana, 1997 —S Sichuan (Daliangshan)

29. Platycerus tieguanzi Imura, 2007 —C Sichuan (Omeishan)

30. Platycerus tangi Imura, 2008 —NC Sichuan (Maoxian)

31. Platycerus cyanidraconis Imura, 2008 —NC Sichuan (Maoxian)

32. Platycerus businskyi Imura, 1996 —Shaanxi (Qinling)

33. Platycerus bashanicus Imura & Tanikado, 1998 —Chongqing (Dabashan) 11

34. Platycerus consimilis consimilis Tanikado & Tabana, 1998 —NC Sichuan (Lixian) Platycerus consimilis phagophilus Imura, 2005 —NE Sichuan (Micangshan)

35. Platycerus yeren Imura, 2008 —Hubei (Shennongjia)

36. Platycerus nagahatai Imura, 2008 —Shaanxi (Qinling)

37. Platycerus ladyae Imura, 2006 —W Sichuan (Erlangshan)

38. Platycerus hiurai Tanikado & Tabana, 1997 —S Sichuan (Daliangshan)

Platycerus hiurai tanikadoi Imura, 2002 —C Sichuan (Xilingxueshan) 39. Platycerus kitawakii Imura & Tanikado, 1998 —Chongqing (Dabashan) 40. Platycerus turnai Imura, 2001 —Hubei (Shennongjia)

Fossil species:

41. Platycerus sepultus Germar, 1837 —Oligocene, Germany 42. Platycerus zherichini Nikolajev, 1990 —Oligocene, Russia

Notes on the checklist:

As there are numerous discrepancies between our checklist and the latest published catalogue of Palaearctic Coleoptera ( Lobl & Smetana 2006) , we provided the following explanations for all of the discrepancies.

1. The synonymic list of American species follows Paulsen (2008).

2. According to Paulsen (2005, 2008), Benesh (1946) erroneously used the name Platycerus piceus Kirby , but that name is not available as it is based on a misidentification of Ceruchus piceus (Weber) . 3. The synonymic list of European and western Asian species follows Lobl & Smetana (2006). 4. According to Imura (1994b, 2006d), Platycerus caerulosus Didier & Seguy is no different from P. caraboides in male genitalia, and it has not been rediscovered in China or even in adjacent areas. The most likely explanation is that the type series was mislabelled from Hunan (Hong-Jang-Hsien). 5. Platycerus chalybaeus MacLeay , Platycerus chlorizans Dalla Torre , and Platycerus coeruleus Dalla Torre were treated as synonyms of Platycerus caucasicus (Parry) by Maes (2008).

6. Platycerus diabolus Dalla Torre and Platycerus nigripes MacLeay were treated as synonyms of Platycerus delagrangei Fairmaire by Maes (2008).

7. Platycerus pseudocaprea Paulus was treated as a valid species by Maes (2008).

8. According to Imura (2007b), the following three taxa: Platycerus acuticollis takakuwai Fujita , Platycerus acuticollis akitai Fujita , and Platycerus acuticollis namedai Fujita constitute a species independent from Platycerus acuticollis Kurosawa , but the formal revision has not been published yet.

9. Platycerus benesi Imura & Bartolozzi was recently synonymized with Platycerus dundai Imura & Bartolozzi by Imura (2005c). This species exhibits great individual variation.

10. Platycerus dundai miyatakei Tanikado & Tabana was originally described as a valid species but was subsequently downgraded by Imura (2005c) as a subspecies of Platycerus dundai Imura & Bartolozzi 11. Platycerus bashanicus Imura & Tanikado was originally described as a subspecies of Platycerus businskyi Imura , but was subsequently given full species rank by Imura (2006b).

Figures

FIGURE 158. Distribution of Platycerus species in China.