Amphidoxa jacquenetta Hutton, 1883

Brook, Fred J., Kennedy, Martyn, King, Tania M., Ridden, Johnathon, Shaw, Matthew D. & Spencer, Hamish G., 2020, Catalogue of New Zealand land, freshwater and estuarine molluscan taxa named by Frederick Wollaston Hutton between 1879 and 1904, Zootaxa 4865 (1), pp. 1-73 : 27-28

Amphidoxa jacquenetta Hutton, 1883

Pl. 3, fig. B

Hutton, 1883 . New Zealand Journal of Science, 1: 476.

Type material. In Canterbury Museum, Christchurch, according to Hutton (1898 – 1900: 7) and Suter (1913: 650), overlooked and reported as missing by Freeman et al. (1997: 36), but one damaged specimen rediscovered in 2017. Lectotype (designated here), CMNZ M277 (dry shell). The molluscan collection at CMNZ has radula fragments mounted on a glass slide with the label details ‘ Amphidoxa jacquenetta, Greymouth, XVI p. 179’, in Hutton’s handwriting (i.e., CMNZ 2017.17.15), which is probably primary type material (see descriptions of radula and jaw by Hutton 1884b: 180 ).

Label details. ‘36. Amphidoxa jacquenetta Hutton , Greymouth’, pillbox label in Hutton’s handwriting.

CMNZ molluscan catalogue details. M277—‘ Flammulina jacquenetta Hutton, Greymouth (1 specimen) (old No. 36) ’.

Type locality. ‘Greymouth (R. Helms)’ ( Hutton 1883g: 476 , 1884b: 180).

Previous illustrations of type material. Radula teeth illustrated by Hutton (1884b: pl. 10, fig. C) probably from type material; Pilsbry (1892 [in 1892–1893]: 76, pl. 22, figs. 70–72—‘drawings of Prof. Hutton’s type, furnished by Mr. Suter’); Suter (1915: pl. 9, figs. 11, a, b—possibly the same shell as illustrated by Pilsbry).

Remarks. Based on one or more specimens collected from Greymouth by Richard Helms. Hutton submitted a description of this species to the Transactions and Proceedings of the New Zealand Institute issue for 1883, but publication was delayed until May 1884 ( Hutton 1884b: 179 ), and was pre-empted by a brief description in an account of a meeting of the Philosophical Institute of Canterbury ( Hutton 1883g: 476 ). From Pilsbry (1893 [in 1893–1895]: 18) and Hedley & Suter (1893: 643) onwards, jacquenetta has generally been assigned to Flammulina Martens, 1873 at subgenus or genus level. However, whereas the protoconch of Flammulina zebra (Le Guillou, 1842) , the type species of this genus, is smooth and lacking in sculpture, that of jacquenetta has a distinctive sculpture of diagonally reticulated riblets with beaded intersections, which indicates that these two species are probably not congeneric. Recorded from Greymouth only by Hutton (1884c: 198) , Suter (1913: 679) and Powell (1979: 314), but examination of material in museum collections indicates that jacquenetta has a sparse distribution in the northwestern South Island (below).

Current taxonomy. Flammulina jacquenetta (Hutton, 1883) — Hedley & Suter (1893: 643), Suter (1913: 678), Powell (1979: 314), Spencer et al. (2009: 215), but the genus level placement requires re-evaluation.

Distribution. New Zealand; northwestern South Island, between Whanganui Inlet and Greymouth (AIM and NMNZ collection records).