Sarax

Rahmadi, Cahyo, Harvey, Mark S. & Kojima, Jun-Ichi, 2010, Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island, Zootaxa 2612, pp. 1-21 : 18

Key to the species of Sarax species from Borneo

1. Anterior margin of carapace rounded; median eye tubercle well-developed or reduced in size; pedipalpal patella with 4 spines, P1 the longest, and subsequent spines decreasing in size from distal to proximal; pedipalpal tarsus with 2 or 3 spines; tibia of leg IV with 17 or 19 trichobothria ..................................................................................... 2

- Anterior margin of carapace slightly rounded, nearly straight ( Fig. 1 ); median eye tubercle reduced in size; pedipalpal patella with 3 spines, about equal in size ( Fig. 4 ); pedipalpal tarsus with 3 spines ( Fig. 5 ); tibia of leg IV with 19 trichobothria ( Fig. 6 ) ........................................................................................................... Sarax cavernicola sp. nov.

2. Pedipalpal patella with 4 major spines on antero-dorsal and 4 or 3 major spines on pedipalpal femur; pedipalpal tarsus with 2 or 3 spines; tibia leg IV with 19 trichobothria ............................................................................................. 3

- Pedipalpal patella with 4 major spines on antero-dorsal and 3 major spines on pedipalpal femur ( Fig. 16 ); pedipalpal tarsus with 2 minute spines ( Fig. 18 ); tibia of leg IV with 17 trichobothria ( Fig. 20 ).................................................... ................................................................................................................................ Sarax sarawakensis ( Thorell 1888)

3. Pedipalpal tarsus with 2 spines; trichobothrium bt on the middle or close to distal fourth basitibial segmenttrichobothrium bc closer to bf than to sbf or about midway between bf and sbf; with or without sexual dimorphism ............. 4

- Pedipalpal tarsus with 3 spines ( Figs. 11 , 13 ); trichobothrium bt close to distal fourth basitibial segment, trichobothrium bc about midway between bf and sbf ( Fig. 12 ); with distinct sexual dimorphism ............ Sarax yayukae sp. nov.

4. With distinct sexual dimorphism; median eye tubercle completely divided into two parts ( Fig. 26 ); trichobothrium bt close to proximal fourth basitibial segment, trichobothrium bc about midway between bf and sbf ( Fig. 29 )................ .................................................................................................................................................... Sarax mardua sp. nov.

- Without sexual dimorphism; median eye tubercle high, black and well-developed, not divided into two parts ( Fig. 21 ); trichobothrium bt on the middle of fourth basitibial segment; trichobothrium bc closer to bf than to sbf ( Fig. 25 ) ................................................................................................................................... Sarax sangkulirangensis sp. nov.

Figures

FIGURES 1 – 7. Sarax cavernicola sp. nov., female holotype: 1. Carapace; 2. Left chelicera, external view; 3. Sternal area of opisthosoma, ventral view; 4. Left pedipalp, antero-dorsal view; 5. Left pedipalpal tarsus; 6. Arrangement of trichobothria on fourth basitibial segment and distibiae of left leg IV; 7. Female gonopods, dorsal view.

FIGURES 16 – 20. Sarax sarawakensis (Thorell 1888), female holotype from Borneo unless stated otherwise: 16. Frontal carapace and right pedipalp; 17. External view of right chelicerae; 18. Pedipalpal tarsus, internal view; 19. Tibial and tarsal segments of leg I showing three most distal tibial segments, and five most proximal tarsal segments; 20. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV, (MZB. Ambl. 149).

FIGURES 8 – 12. Sarax yayukae sp. nov., male holotype, unless stated otherwise: 8. Frontal area of carapace, male paratype from Liang Hajuq (MZB. Ambl. 148); 9. Left chelicerae, external view; 10. Sternal area of opisthosoma, ventral view; 11, Left pedipalp, antero-dorsal view; 12. Arrangement of trichobothria of fourth basitibial segment and distibiae of left leg IV.

FIGURES 13 – 15. Sarax yayukae sp. nov.: 13. Left pedipalp, antero-dorsal view, female paratype (MZB. Ambl. 057); 14. Female genitalia, paratype (MZB. Ambl. 057); 15. Pedipalp and anterior margin of carapace, juvenile paratype (MZB. Ambl. 059).

FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12).

FIGURES 21 – 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150).