Incoltorrida Steffan, 1973

Perkins, Philip D. & Bergsten, Johannes, 2019, New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae), Zootaxa 4657 (1), pp. 57-96 : 64

Incoltorrida Steffan

Incoltorrida Steffan, 1973: 634 . Type species Incoltorrida madagassica Steffan, 1973 (by original designation)

The following comments are given by Steffan (1973) as the differential diagnosis of Incoltorrida : “The members of Incoltorrida n. gen. differ from the members of Torridincola Steffan 1964 , through the following characters: on average more robust body size and more pronounced sculpturing of the elytra; the proximally narrowed keel on frons and vertex; the length proportions of the tarsomeres (mesotarsi T4 approximately 1.5 times longer than T3, compared to twice as long in Torridincola ); The absence of parameres and the prescence of a thorned saccus membranaceus in the male, and the lateral thorns on the styli of the female genital organ.”

All of these diagnostic characters hold true for the new species described herein, except the presence in all species of a single seta on each side near the base of the male genitalia (e.g., Figs. 19, 20 ); these setae confirm that the basal lobes they are attached to are the homologues of the parameres present in members of Torridincola . Because of distortion, these setae are not visible on the permanent microslide mount of the holotype of I. madagassica , the only male available to Steffan at the time of the original description.

Figures

FIGURES 19–20. (19) Incoltorrida madagassica, male genitalia of non-type; (20) Incoltorrida benesculpta, male genitalia of holotype.