Incoltorrida marojejy, Perkins & Bergsten, 2019

Perkins, Philip D. & Bergsten, Johannes, 2019, New Myxophagan water beetles from Madagascar (Coleoptera: Torridincolidae, Hydroscaphidae), Zootaxa 4657 (1), pp. 57-96 : 73-75

Incoltorrida marojejy , new species

Figs. 9 (habitus); 22 (genitalia); 24 (elytron); 37 (map); 41, 44–47 (habitat)

Type Material. Holotype (male): Antsiranana, 14.4345S 49.7606E Sava Marojejy National Park , Camp II, wide river forming hygropetrick rocks and rockpools, elev. 710 m, 9 xi 2014, J. Bergsten, R. Bukontaite, J.H. Randriami- haja, T. Ranarilalatiana & S. Holmgren ( MAD 14-54 ) ( NHRS) . Paratypes: Antsiranana: Sava: Marojejy National Park : midaltitude rainforest: MAD 18-24 : upstream on stream by camp II, traversing from Taktajania trail, 14.4369S, 49.7603E, 820m, 09.II.2018, J.Bergsten & T.Ranarilalatiana (98 NHRS, MCZ, BMNH, PBZT / MBC) .

Differential Diagnosis. Pronotal sculpture similar to I. magna ; both species lack ridges on each side near the base. Differing therefrom by the smaller size (ca. 2.19 vs. 2.62 mm), the presence of transverse grooves on the meta- ventrite tabella, the shape of the frontoclypeal shield, and the larger serial elytral punctures. The aedeagi of the two species differ markedly ( Figs. 21, 22 ).

Description. Size: holotype (length/width, mm): body (length to elytral apices) 2.19/1.26; head width 0.41; pronotum 0.47/0.89; elytra 1.40/1.26. Dorsum brown to reddish brown, venter reddish brown, legs reddish brown except femoral-tibial articulations and tips of tarsi black.

Sides of frontoclypeal shield slightly arcuate toward midline, such that apical part is slightly wider than width at midlength, and slightly narrower than posterior of frons.

Short oblique carinae on anterior 1/3 of pronotum well developed, no indication of midlongitudinal carina, and area posterior to carinae is transversely rounded and shows no indication of ridges or depressions.

Elytron with eight strong costae; costae #6 and #7 interrupted by distinct transverse ridge that connects #5 and #8. Serial punctures large and deep.

Posterior 1/2 of metaventral tabella with distinctive, closely spaced transverse grooves; midlongitudinal groove in posterior 1/2 of metaventral tabella gradually widens from posterior to anterior.

Midlongitudinal carina of first abdominal ventrite strong and extends length of ventrite.

Male genitalia in lateral view distinctively thickened in basal 1/2, apical tip abruptly distinctively narrowed and spinose ( Fig. 22 ).

Etymology. Named in reference to the type locality.

Remarks. Four larvae were collected in association with adults at site MAD 18-24.

Figures

FIGURE 9. Incoltorrida marojejy, habitus views of holotype.

FIGURES 21–22. Incoltorrida, male genitalia of holotypes. (21) I. magna; (22) I. marojejy.