Claustropyga glacialis, Rudzinski & Ševčík, 2019

Rudzinski, Hans-Georg & Ševčík, Jan, 2019, Czech and Slovak Claustropyga (Diptera: Sciaridae), with the description of C. glacialis sp. nov. and notes on morphological variation in some congeners, Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (2), pp. 529-535 : 530

Claustropyga glacialis sp. nov.

( Figs 1–8 )

Type material. HOLOTYPE: J, CZECH REPUBLIC: MORAVIA & SILE- SIA: Hrubý Jeseník Mts, Praděd National Nature Reserve, Velká kotlina glacial cirque, 9.–26.vi.2006, J. Roháček & J. Ševčík leg. (Malaise trap). Prep. - Nr.: CZ/784 ( SMOC) . PARATYPE: J, CZECH REPUBLIC: BOHE- MIA: Šumava Mts, Horská Kvilda, montane peat-bog, 20.v.–17.vi.1999, M. Barták & J. Roháček leg.(Malaise trap). Prep.-Nr.: CZ/843 ( NMPC).

Description. Male. Body length: 3 mm. Head. Eye bridge 3 facets wide. Face (prefrons) with 9 setae. Clypeus with 0–1 seta. Maxillary palpus ( Fig. 1 ) short, brownish, with 3 segments. Palpomere 1 with 1 long dorsolateral seta and diffuse patch of dorsal sensillae. Length of palpomeres = 70:50:60 μm. Antenna ( Fig. 2 ) brown. Body of antennal flagellomere 4: 2.1× as long as wide. Length of flagellomere necks: 0.16× length of flagellomere 4. Longest setae shorter than width of flagellomeres. Thorax dark brown. Scutum with dark and strong lateral and dorsocentral setae. Scutellum with 2 long apical marginal setae. Anterior pronotum with 2 setae. Episternum 1 with 6 setae. Postpronotum and other pleural tergites non-setose. Coxae pale brownish, with dark setae. Haltere dark brown; H-K-index = 2.5. Wing pale brownish. Anal lobe well developed. C/w = 0.70. R1/R = 1.07. R-m/bM = 0.81. R-m distal with 1–2 dorsal setae; bM non-setose. Base of CuA very short: 0.16× length of bM. Wing length = 3.1 mm. Legs yellowish brown. T1 without ventral spines. Front tibial organ ( Fig. 3 ) with strong setae in 3–5 irregular rows; proximally without distinct bordering. T 3 in apical half with line of fine dorsal spines. Abdomen brown, concolorous with thorax. Tergal setae strong, dense and dark. Terminalia. Tergite 9 ( Fig. 4 ) apically rounded; with dense and short setosity. Hypopygium ( Fig. 5 ) dark brown.Intercoxal area produced, with dense and short setosity. Gonostylus ( Figs 6–8 ) swollen; medially deeply concave, like splitting into ventral and dorsal lobe. Ventral lobe ( Fig. 7 ) with 4 megasetae in apical-subapical position and long setosity. Dorsal lobe ( Fig. 8 ) with short and strong apical tooth, and 4 megasetae; 2 megasetae above apical tooth, 1 megaseta below tooth, and 1 megasetae subbasal on inner side of gonostylus.Tegmen pyramidal; lateral shoulders with line of fine marginal teeth. Aedeagus strongly developed. Aedeagal membrane with patch of fine teeth.

Female unknown.

Differential diagnosis. Regarding the special structure of the gonostylus, Claustropyga glacialis sp. nov. is similar to C. lobigera Hippa & Vilkamaa, 2016 and C. mirifica Vilkamaa & Hippa, 2007 . Both species differ from C. glacialis sp. nov. in having a dorsal pair of long narrow lobes on the tegmen. Claustropyga mirifica also differs from C. glacialis sp. nov. in having no apical tooth on the gonostylus. Claustropyga lobigera differs from the new species in having only 3 apical-subapical megasetae besides the short apical tooth.

Etymology. The species epithet is the Latin adjective glacialis (- is, - e), meaning icy, frozen or glacial. The name is given in reference to its habitat.

Habitat. The type specimens were collected in a glacial cirque and montane peat-bog, respectively, suggesting that the new species may be considered a glacial relict in Central Europe.

Figures

Figs 1–8. Claustropyga glacialis sp. nov. (holotype), 1 – maxillary palpus; 2 – antennal flagellomere 4; 3 – front tibial organ; 4 – 9th tergite; 5 – hypopygium, ventral view; 6 – gonostylus, ventral view; 7 – ventral lobe of gonostylus, inner side; 8 – dorsal lobe of gonostylus, inner side.