Clada (Clada) madagascarensis, Zahradnik, Petr & ryzna, Milos, 2018

Zahradnik, Petr & ryzna, Milos, 2018, Nine new species of Clada from Madagascar (Coleoptera, Ptinidae), ZooKeys 806, pp. 121-140 : 121

Clada (Clada) madagascarensis sp. n. Figs 5, 14, 23a, b, 32

Type material.

Holotype male: Madagascar, Mahajanga distr., Ampatika env., 17.-20.xi.1995, J. Stolarczyk lgt. (FGMRI). Paratypes (21): 4 males and 1 female, the same data as holotype (FGMRI); 10 males, Madagascar, Morondava distr., Kirindy, 23.-25.xi.1997, J. Stolarczyk lgt. (FGMRI 4 ex., LBVC 2 ex., MTDC 2 ex., NHMUK 2 ex.); 1 male: Madagascar, Mahajanga distr., Ambodimanga env., 14.-16.xi.1995, J. Stolarczyk lgt. (FGMRI); 2 males: Madagascar, Morondava distr., Maronfandilia, 4.v.1995, J. Stolarczyk lgt. (FGMRI); 2 male: Madagascar, Mahajanga prov., Mahajamba riv., Ampatika env., 17.-19.xi.1995, I. Jeniš lgt. (FGMRI); 1 female: Madagascar, Toliara env., 23.-27.xi.1996, J. Stolarczyk lgt. (FGMRI).

Differential diagnosis.

This species is similar to C. (C.) humeralis Pic, 1926, but differs by the lighter colour of the elytra and missing lighter humeri. Both sexes have pectinate antennae, while the female of C. (C.) humeralis Pic, 1926 has serrate antennae. Fully differs by shape of the aedeagus. For differences from other Madagascan species, see key.

Description.

Male (holotype). Elongate-elliptical, transversally convex. Body length 6.0 mm, maximum width 2.5 mm (Figure 5). Ratio length:width of elytra 1.6. Whole body brown, only antennae, palp slightly lighter and pronotum slightly darker. Pubescence white.

Head shiny, with double punctation - one coarse, dense, umbilicate, distance between punctures approximately the same as their diameter, sometimes almost touching; other one is very fine, punctures almost touching. Pubescence recumbent or semi-erect, long, inclined mostly forwards. Clypeus with shallow transverse depression. Eyes large, globular with short erect sparse pubescence. Frons twice as wide as diameters of the eye, from dorsal view. Antennae consisting of eleven antennomeres, 4th to 10thpectinate (Figure 23a). 1st antennomere robust, twice as long as wide; 2nd smallest, only 1/3 long as 1st, as wide as long, same width as the 1st. 3rd serrate, as long as wide; 4th to 8th twice long as wide; 9th 1.7 times as wide as long; 10th 1.5 times as wide as long. Apical antennomere longest, oblong oval, 6.6 times as long as wide. All antennomeres with short recumbent pubescence, only 1st with a few long semi-erect setae. Apical maxillary palpomere long, spindle shaped.

Pronotum convex, matt-shiny, transverse (ratio length:width of pronotum 0.7); widest in middle. Base of pronotum finely bordered. Middle of pronotum at base with blunt small swelling, posteriorly slightly sharpened. Surface of pronotum with double punctation - one coarse, dense, umbilicate, distance between punctures approximately the same as their diameter; other one is very fine, punctures almost touching. Pubescence long, sparse, recumbent, inclined more or less to middle of pronotum.

Scutellum large, longitudinally trapezoidal, narrow, almost as long as wide, densely recumbent pubescence, inclined backwards, surface almost invisible.

Elytra oval, transversally convex, shining, with distinct humeri. Each elytron with five very fine costae, more distinct on second half of elytron. Surface of elytra irregular punctated, puncture coarse, dense, umbilicate. Pubescence relatively sparse, recumbent, inclined backwards. Posterior margin of each elytron with approximately 25 very small teeth.

Legs stout, with short and dense recumbent pubescence. All tarsi robust, same length as tibia. 1st metatarsomere as long as 2nd and 3rd together, same width, slightly emarginate on top, 4th slightly shorter than previous, more emarginate, almost to 2/3 of their length. 5th is same length as 3rd and 4th together, rectangular, wider on the top, with two large claws, without teeth.

For aedeagus see Figure 32.

Female. Antennae less pectinate than in male (damaged, only six antennomeres remain - Figure 23b). Body length 5.7 mm, maximum width 2.1 mm.

Variability.

Body length 5.7-7.1 mm, maximum width 2.1-2.8 mm.

Name derivation.

Latin adjective, referring to the occurrence of the new species in Madagascar.

Biology.

Unknown.

Distribution.

This species is found in the western part of Madagascar (Figure 14).