Sarcophaga (Heteronychia) lejlekensis, Whitmore, Daniel, 2011

Whitmore, Daniel, 2011, New taxonomic and nomenclatural data on Sarcophaga (Heteronychia) (Diptera: Sarcophagidae), with description of six new species, Zootaxa 2778, pp. 1-57 : 8-10

Sarcophaga (Heteronychia) lejlekensis sp. nov.

( Figs 12–17 )

Type material. Holotype 3: [transliterated from Cyrillic] Saj Sagul / Kirg. [= Kyrgyzstan] 30 km / s-v Ljajljaka [= N-E of Lejlek] / L. Primykina / [vertically on left side of label] 3.VI.64 [1964] // [red label] HOLOTYPE 3 / Sarcophaga / ( Heteronychia ) / lejlekensis sp. n. / det. D. Whitmore 2010 ( ZIN). Paratypes: 1 3: same locality and date as holotype ( ZIN); 1 3: Kyrgyzstan, 30 km N-E of Lejlek, 2.VI.64, leg. Primykina ( ZIN); 1 3: Kyrgyzstan, 20 km N of Lejlek, 13.9.68, leg. Primykina ( ZIN); 1 3: Kyrgyzstan, Osh province, Sary-Tash, 2– 7.IX.971, leg. Primykina ( MMBC as “ Heteronychia peckae Verves , det. Verves”).

Diagnosis (3). A medium-sized to large species of Heteronychia with a wide frons; lower facial margin slightly protruding in lateral view; scutellum with a pair of apical setae; wing vein R1 bare on dorsal surface; costal spine very short or not developed; abdomen with dense grey microtrichosity forming a maculate pattern in posterior view; abdominal tergite 3 without median marginal setae; epandrium red-orange; cercus with a low, rounded dorsal subapical hump; distiphallus: apical process of harpes flattened, tapering and directed apico-ventrally; juxta short, with triangular lateral processes.

Description (3). Length. 10.0−12.5mm. Colour. Ground colour black, with dense light-grey microtrichosity on parafacials, fronto-orbital plate, thorax and abdomen. Thorax with three longitudinal dark vittae; microtrichosity of abdomen forming typical chequered pattern changing with the incidence of light, lateral black markings on tergites 3−4 reduced to an anterior spot when viewed posteriorly. Protandrial segment with a large rounded patch of microtrichosity near margin; epandrium red-orange. Cercus black; surstylus, phallus and gonites dark brown. Head ( Fig. 12 ). Arista thickened on approximately basal 1/3–2/5. Postpedicel ca. 1.5 times as long as pedicel. Frons at its narrowest point about 0.6–0.7 times the width of an eye in dorsal view. Lateral vertical setae short, but visibly stronger than postocular setae. Parafacial at its narrowest point about 0.3–0.4 times eye width. Lower facial margin slightly protruding in lateral view below vibrissa. Gena in profile about 0.4–0.5 times height of eye; postgena entirely covered with white setulae. Occipital setulae white below the first two rows. Thorax. Scutum with 3 (unarranged) + 1 acrostichal, 4 + 3 dorsocentral, 2 posthumeral (outer one often weak), 1 presutural, 4 notopleural, 2–3 intraalar and 3 (sometimes two additional shorter ones present) supraalar setae. Scutellum with a pair of strong apical setae; discal setae situated far from margin. Legs. Mid femur without a subapical posteroventral comb. Mid tibia with 2–3 anterodorsal, 2–3 posterodorsal, 1 dorsal and 1 anteroventral setae. Hind trochanter with a ventral brush of tightly-spaced, spine-like setae. Hind femur with a few, weak (rarely strong) anteroventral setae in addition to subapical one. Hind tibia with 1 anteroventral seta; hind tibia with several long, wavy setulae on posteroventral surface. Wing. Costal spine very short or not developed. Vein R1 bare on dorsal surface. Second costal section visibly longer (about 1.3x) than fourth costal section. Abdomen. Tergite 3 without median marginal setae. Terminalia. Setae on sternite 5 thickened and shortened, forming a tight brush-like structure; marginal setae visibly longer. Protandrial segment with a row of setulae along posterior margin. Epandrium with an elongated ventral margin. Cercus ( Fig. 13 ) with a low, rounded dorsal subapical hump and more or less uniformly covered in setae; tip downcurved, hooked, with a convex dorsal surface. Surstylus ( Fig. 13 ) subtriangular. Pregonite ( Fig. 14 ) short, with a sharply pointed, hooked tip, and with several long setae on entire dorsal surface. Distiphallus ( Fig. 15 ): proximal part of harpes rounded in lateral view, with a concave inner surface and continuous with distal part; apical process flattened, tapering and directed apico-ventrally; juxta short, with triangular lateral processes, trilobate in apical view ( Fig. 16 ) with two blunt lateral processes and a subtrapezoidal median process; vesica ( Fig. 17 ) well developed, short, rounded in cross section, with pointed corners.

Female unknown.

Distribution. Kyrgyzstan.

Etymology. A Latin adjective referring to the Lejlek region in Kyrgyzstan, type locality of the species.

Differential diagnosis. Sarcophaga (Heteronychia) lejlekensis sp. nov. has been confused, in museum collections (ZIN, MMBC), with S. (H.) peckae (Verves, 1977) , also described from Kyrgyzstan (Tien Shan) and which is in reality a junior synonym of S. (H.) shnitnikovi ( Rohdendorf, 1937) (see below). The new species differs from S. (H.) shnitnikovi by the denser microtrichosity of the abdomen (appearing ‘maculate’ when viewed posteriorly) and by the shape of the cercus and distiphallus (compare with Figs 74–75 ).

Figures

FIGURES 12 – 17. Sarcophaga (Heteronychia) lejlekensis sp. nov. 12. Head, holotype, scale bar 1 mm. 13 – 17. Paratype from Kyrgyzstan, Sary-Tash (MMBC). 13. Cercus and surstylus, scale bar 0.25 mm. 14. Gonites, scale bar 0.1 mm. 15. Distiphallus (lateral), scale bar 0.1 mm. 16. Juxta (apical), scale bar 0.05 mm. 17. Vesica, scale bar 0.05 mm.

FIGURES 66 – 77. 66 – 68. Sarcophaga (Heteronychia) pseudobenaci (Baranov). 66. Cercus and surstylus, specimen from Greece, Hortiatis (CNBFVR), scale bar 0.15 mm. 67 – 68. Specimen from Greece, Stávros (ZMUC), scale bars 0.1 mm. 67. Distiphallus (lateral). 68. Juxta (dorsal). 69 – 71. Sarcophaga (Heteronychia) benaci Böttcher, specimens from Croatia, Krk (NHRS). 69. Cercus and surstylus, scale bar 0.1 mm. 70. Distiphallus (lateral), scale bar 0.05 mm. 71. Juxta (dorsal), scale bar 0.05 mm. 72 – 73. Sarcophaga (Heteronychia) infantilis Böttcher, specimen from Austria, Steinernes Meer (NHRS). 72. Cercus and surstylus, scale bar 0.1 mm. 73. Distiphallus (lateral), 0.05 mm. 74 – 75. Sarcophaga (Heteronychia) shnitnikovi (Rohdendorf), specimen from Kyrgyzstan, Kaindy (ZIN). 74. Cercus and surstylus, scale bar 0.25 mm. 75. Distiphallus (lateral), scale bar 0.1 mm. 76 – 77. Sarcophaga (Heteronychia) vagans Meigen, specimen from Austria, Niedere Tauern (ZMUC), scale bars 0.25 mm. 76. Phallus and gonites. 77. Cercus and surstylus.