Sarcophaga (Heteronychia) ancilla Rondani, 1865

Whitmore, Daniel, 2011, New taxonomic and nomenclatural data on Sarcophaga (Heteronychia) (Diptera: Sarcophagidae), with description of six new species, Zootaxa 2778, pp. 1-57 : 17-19

Sarcophaga (Heteronychia) ancilla Rondani, 1865

( Figs 45–46 )

Sarcophaga ancilla Rondani, 1865: 226

Heteronychia iubita Lehrer, 1999: 410 , syn. nov.

Spatulapica lucentina Lehrer & Martínez-Sanchez, 2001: 3 , syn. nov.

Type material examined. Sarcophaga ancilla : Holotype 3: 1029 // Lectotypus / Sarcophaga / ancilla Rond. // Heteronychia / ancilla / det. F. Mihalyi // HOLOTYPE 3 / Sarcophaga / ancilla Rond. / T. Pape det. 1968 [sic! = 1986] (see Pape 1988).

Heteronychia iubita : Holotype 3: 10.V.1977 / leg. Kugler / FRANCE / Provence / Lagnes // Heteronychia 3 / iubita sp. n. / HOLOTYPUS / Det. Dr. A.Z. Lehrer / 1998 // Heteronychia 3 / iubita sp. n. / HOLOTYPUS / Det. Dr. A.Z. Lehrer /1998 ( TAU); terminalia dissected and mounted on three separate slides, labelled as follows: Heteronychia 3 / iubita n. sp. / Det. Dr. A.Z. LEHRER // 3 HOLOTYPUS / Sternite V ( TAU); Heteronychia 3 / iubita n. sp. / Det. Dr. A.Z. LEHRER // 3 HOLOTYPUS / Cerci & paralobi ( TAU); Heteronychia 3 / iubita n. sp. / Det. Dr. A.Z. LEHRER // 3 HOLOTYPUS / Phallosome ( TAU).

Additional material examined. France: Bouches-du-Rhône, Chaines-des-Alpilles, 21.IV.1994, R. Blackith leg., 6 3 ( NHRS); Vingrau (66), V.2004, R. Richet leg., 1 3 ( NHRS). Italy: 2 3: 1020 ( MZUF) [these are paralectotypes of Pierretia (Heteronychia) rondaniana Rohdendorf, 1937 designated by Pape (1988: 17), and were both misidentified by Pape: one as S. (H.) depressifrons Zetterstedt , the other as S. (H.) bezziana Böttcher (= infantilis Böttcher ) (see Pape 1988)]; Latium, Rieti prov., Amatrice, 7.VII.1897, 2 3 ( SMF, Böttcher collection); Lombardy, Pavia, 10.VI.1890, 1 3 ( SMF, Böttcher collection); Tuscany, Firenze, 10–18.V.1986, T. Pape leg., 3 3 ( ZMUC). Spain: Lerida, Sierra de Boumort, 24.VI. 1982, 900m, Andersen, Lyneborg & Michelsen leg., 3 3 ( ZMUC); Lerida, Tremp, 1–12.VII.1981, V. Michelsen, 1 3 ( ZMUC). Switzerland: Valais, Leuk-Pfynwald, 630m, 25.V.1997, B. Merz leg., 1 3 ( NHRS).

Remarks. Pape (2004a) synonymized Heteronychia iubita with Sarcophaga (H.) depressifrons Zetterstedt, 1845 . Direct examination of the holotype, and the original description and illustrations ( Lehrer 1999: 410 + fig. 1) allow me to formalize that H. iubita is a junior synonym of S. (H.) ancilla .

Spatulapica lucentina was described on a single specimen from Sierra Salinas (Alicante, Spain), deposited at DCAA ( Lehrer & Martínez-Sanchez (2001: 221). I did not examine the holotype, but the original description and illustrations ( Lehrer and Martínez-Sanchez 2001: 221, figs 1–5) correspond exactly to Sarcophaga (Heteronychia) ancilla . Sarcophaga (H.) ancilla is a small species (3–4mm) with a shiny black epandrium, showing only slight variation in the shape of the cercus and distiphallus; the apical process of harpes is relatively long ( Fig. 45 ), and the tip of juxta is narrow (less than 1/4 of width of juxta in apical view) ( Fig. 46 ). Slightly larger specimens with a reddish-black to entirely red epandrium, a much shorter apical process of harpes and a much wider tip of juxta (about 1/3 of width of juxta in apical view), which have often been referred to as S. ancilla in the literature (e.g., Rohdendorf 1937: 356, figs 483–484; Povolný 1996: 106, fig. 19b), belong to Sarcophaga (H.) belanovskyi (Verves, 1973) (see below). Due to the confusion of the two species over a long period of time, the distribution of S. (H.) ancilla (see Pape 1996, 2004a) is in need of revision, and may even be restricted to south-western Europe. Illustrations of “ Heteronychia ancilla ” by Povolný (1996: 106, fig. 19a) and Povolný and Verves (1997: 165, fig. 177) refer to S. (H.) lederbergi ( Lehrer, 1995a) .

Both paralectotypes of Pierretia (Heteronychia) rondaniana designated by Pape (1988: 17) belong to S. (H.) ancilla (see above).

Figures

FIGURES 42 – 53. 42. Sarcophaga (Heteronychia) thirionae (Lehrer), juxta (dorsal); specimen from Greece, Olympos (MMBC), scale bar 0.1 mm. 43 – 44. Sarcophaga (Heteronychia) villeneuveana (Enderlein), specimen from France, Maury (CNBFVR). 43. Juxta (dorsal), scale bar 0.05 mm. 44. Distiphallus (ventral), scale bar 0.1 mm. 45 – 46. Sarcophaga (Heteronychia) ancilla Rondani, scale bars 0.05 mm. 45. Distiphallus (lateral), specimen from France, Bouches-du-Rhône (NHRS). 46. Distiphallus (apical), specimen from Spain, Sierra de Boumort (ZMUC). 47 – 48. Sarcophaga (Heteronychia) belanovskyi (Verves), specimen from Italy, Castelporziano (MZUR), scale bars 0.05 mm. 47. Distiphallus (lateral). 48. Distiphallus (apical). 49 – 50. Sarcophaga (Heteronychia) chiquita (Peris et al.). 49. Cercus and surstylus, paratype from Spain, Pina de Ebro (UCME), scale bar 0.1 mm. 50. Juxta (lateral), holotype of Heteronychia morenita Peris et al., scale bar 0.05 mm. 51 – 53. Sarcophaga (Heteronychia) croca Pape. 51. Cercus and surstylus, specimen from Croatia, Hvar (NHRS), scale bar 0.25 mm. 52 – 53. Specimen from Greece, Skotina (MMBC). 52. Pregonite, scale bar 0.05 mm. 53. Distiphallus (lateral), scale bar 0.1 mm.