Sarcophaga (Heteronychia) nanula ( Povolný, 1999 ), Povolny, 1999

Whitmore, Daniel, 2011, New taxonomic and nomenclatural data on Sarcophaga (Heteronychia) (Diptera: Sarcophagidae), with description of six new species, Zootaxa 2778, pp. 1-57 : 44

Sarcophaga (Heteronychia) nanula ( Povolný, 1999) , revived stat.

( Fig. 64 )

Heteronychia rohdendorfi ssp. nanula Povolný, 1999: 11 .

Type material examined. Heteronychia rohdendorfi ssp. nanula : Holotype 3: SICILIA / [Palermo prov.] Pizzo Cane-Savochetto / 882 m a. s. l. / June 2, 1998 // Spatulapica / gr. rohdendorfi / det. Povolný [/] gr. rohdendorfi / 2.VI.98 / Pizzo Cane // [red label] Holotypus / det. Povolný [/] Heteronychia / rohdendorfi / nanula // TRAN- SCRIPTIO / Heteronychia / rohdendorfi nanula ssp. n. / det. Povolný 1999 // Invent. č. / 5593 / Ent. / Mor. muzeum, Brno ( MMBC) [holotype with last abdominal segments and terminalia removed and placed in glycerin in a microvial pinned beneath the source specimen]. Paratypes: 3 3: one with locality label 2.VI.98 / Pizzo Cane, two with red “ Paratypus ” label but no locality label ( MMBC).

Additional material examined. France (new country record): Bouray, 1.VI.1916, 1 3 ( IRSNB). Italy: Sicily, Palermo prov.: Bosco della Ficuzza, nr. Alpe Cucco, 776m, 23.VI.2005, D. Whitmore et al. leg., 1 3 (CNBFVR); Casina, Valle del Corvo, V.2001, D. Povolný leg., 2 3 ( MMBC); same locality, V–VI.2002, D. Povolný leg., 1 3 ( MMBC); Siracusa prov., Sortino, Necropoli di Pantalica, 15.V.1968, S. Langemark leg., 1 3 ( ZMUC). Macedonia (new country record): Skopje, 25.VI.1927, N. Baranov leg., 1 3 ( USNM).

Remarks. Povolný (1999) described this taxon as a subspecies of Heteronychia rohdendorfi (= S. (H.) lederbergi ), and Pape (2004a) subsequently synonymized it under S. (H.) lederbergi . After examining the holotype, paratypes, and a few additional specimens, I consider nanula as a valid species of Sarcophaga (Heteronychia) , possibly more closely related to S. (H.) depressifrons than to S. (H.) lederbergi . It differs from the former by the denser microtrichosity of the thorax and abdomen (which makes it appear of a lighter grey colour), the white occipital setulae behind the first two rows, the light red epandrium (usually black/dark brown in S. depressifrons ) and the wider tip of juxta in dorsal/apical view (compare Figs 64–65 ). It differs from S. lederbergi by the narrower and shorter lateral styli and the narrower juxtal tip in dorsal/apical view.

Figures

FIGURES 54 – 65. 54 – 56. Sarcophaga (Heteronychia) enderleini Jacentkovský. 54. Cercus and surstylus, specimen from Italy, Abetina di Rosello (CNBFVR), scale bar 0.25 mm. 55. Distiphallus (lateral), type specimen of Heteronychia (Eupierretia) macedonica Povolný from Greece, St. Panteleimon (MMBC), scale bar 0.1 mm. 56. Juxta (apical), specimen from Italy, Borgo Capanne (NHRS), scale bar 0.05 mm. 57. Sarcophaga (Heteronychia) ferox Villeneuve, distiphallus (half ventral, arrow showing vesica); paratype of Heteronychia (Asceloctis) perplexa Peris et al. from Spain, Pego, scale bar 0.15 mm. 58. Sarcophaga (Heteronychia) giganta Pape, distiphallus (lateral); specimen from Croatia, Biokovo (NHRS), scale bar 0.25 mm. 59. Sarcophaga (Heteronychia) kataphygionis (Povolný), distiphallus (lateral); paratype from Greece, Olympos (MMBC), scale bar 0.1 mm. 60. Sarcophaga (Heteronychia) sp., distiphallus (lateral); specimen from Ukraine, Andriyivka (YVC), scale bar 0.1 mm. 61. Sarcophaga (Heteronychia) kerteszi Villeneuve, terminalia; specimen from Lebanon, Mt. Barukh (TAU, as “ Heteronychia dayani det. Lehrer ”), scale bar 0.25 mm. 62 – 63. Sarcophaga (Heteronychia) minima Rondani, specimen from Israel, Hameshar (TAU, as “ Ctenodasypygia fertoni det. Lehrer ”). 62. Cercus and surstylus, scale bar 0.15 mm (arrow showing subapical hump of cercus). 63. Distiphallus (lateral), scale bar 0.1 mm (arrow showing protruding lateral membranous process of juxta). 64. Sarcophaga (Heteronychia) nanula (Povolný), distiphallus (apical); specimen from Sicily, Sortino (ZMUC), scale bar 0.05 mm. 65. Sarcophaga (Heteronychia) depressifrons Zetterstedt, distiphallus (apical); specimen from Switzerland, Zürichberg (NHRS), scale bar 0.05 mm.