Sarcophaga (Heteronychia) vagans Meigen, 1826

Whitmore, Daniel, 2011, New taxonomic and nomenclatural data on Sarcophaga (Heteronychia) (Diptera: Sarcophagidae), with description of six new species, Zootaxa 2778, pp. 1-57 : 50-51

Sarcophaga (Heteronychia) vagans Meigen, 1826

( Figs 76–77 )

Sarcophaga vagans Meigen, 1826: 26 .

Sarcophaga nepos Rondani, 1860: 390 .

Sarcophaga frenata Pandellé, 1896: 182 .

Sarcophaga bajkalensis Rohdendorf, 1925: 56 .

Hartigia anastrenua Baranov, 1942: 621 , syn. nov. Heteronychia (Eupierretia) vagans ssp. sachalinensis Verves, 1978b: 185.

Type material examined. Sarcophaga vagans : Syntype Ƥ: [round label] 2090 / 40 // Sarcophaga / vagans / Ƥ // Holotype Ƥ / Sarcophaga / vagans Meig. ( MNHN) [terminalia dissected (by R. Richet) and placed in Canada balsam in a small slide pinned beneath the source specimen; first abdominal segments glued to the upper surface of the square of material used to enclose the slide].

Remarks. Meigen (1826) described Sarcophaga vagans on both sexes, as confirmed by the original identification label pinned beneath the syntype found in MNHN, as well as by the relatively low position of the female on the pin. The type series therefore consisted of at least two specimens (in all probability just the pair), and Richet’s labelling of the above female of Sarcophaga vagans as “ holotype ” is incorrect. The female, with a pentagonal vaginal plate, corresponds to S. (H.) vagans as treated by Rohdendorf (1937, as “ Pierretia (Eupierretia) frenata ”), Pape (1987), Povolný and Verves (1997) and Richet et al. (in press) ( Figs 76–77 ).

Sarcophaga nepos : Lectotype 3 (designated by Pape 1988): 1033 // = cruentata / sec. Typ. Pand. / det. Böttch- er // LECTOTYPE 3 / Sarcophaga / nepos Rond. / T. Pape det. // Heteronychia / vagans (Meig.) / T. Pape det. 1986 ( MZUF).

Sarcophaga frenata : Lectotype 3 (designated by Pape 2004b): 3 / 2217 // LECTOTYPE 3 / Sarcophaga / frenata Pnd. / T. Pape det. 2002 (MNHN) (see Pape 2004b).

Hartigia anastrenua : Holotype 3: Croatia / Zagreb / 22.V.1931. // N. Baranov / Coll. 1960 // [red label] Type No. / 65730 / U.S. N.M. // Hartigia / anastrenua / sp. n. / N. Baranov // HOLOTYPE / Hartigia / anastrenua / BAR. /det. / Sabrosky // Sarcophaga / ( Heteronychia ) / vagans / Meigen, 1826 / det. D. Whitmore 2008 (USNM) [holotype with most of abdomen and terminalia detached and placed in glycerin in a microvial pinned beneath the source specimen].

Heteronychia (Eupierretia) vagans ssp. sachalinensis: Holotype 3: [transliterated from Cyrillic] Yuzhn.- Sachalinsk / 23.5.1957 / Violovich // Holotypus Hetero - / nychia (Eupierretia) va - / gans sachalinensis Verves // Sarcophaga / vagans / det. D. Whitmore 2007 (ZIN) [terminalia detached from abdomen and placed in glycerin in a microvial pinned beneath the source specimen].

Additional material examined. Austria: Hochkar, 4.VIII.1995, D. Povolný leg., 5 3 ( MMBC); Muggendorf, VII. 1999, 1 3 ( ZMUC); Niedere Tauern, Solker pass, 1900m, 7–8.VIII.1995, M. Barták leg., 1 3 ( ZMUC); Schladminger Tauern, Donnersbachwald, 1600m, 5–7.VIII.1995, M. Barták leg., 2 3 ( ZUMC); Seefeld, 19.VII.1960, A. Collart leg., 2 3 ( IRSNB); Staatz, 9.V.1996, D. Povolný leg., 1 3 ( MMBC). Belgium: Fonds de Forêt, 15.VIII.1946, A. Collart leg., 1 3 ( IRSNB). Croatia: Zagreb, N. Baranov leg. several 3 ( USNM); same locality, 18.IX.2003, D. Povolný leg., 1 3 ( MMBC). Czech Republic: Beskydy, Horní Lomná, nr. Mionši, 26– 28.VII.1996, M. Barták leg., 1 3 ( ZMUC); Beskydy, Murinkovy vrch, 950m, M. Barták leg., 10.VI.1995, M. Barták leg., 2 3 ( ZMUC); Brno, Rajhrad, 21.V.1941, 1 3 ( MMBC); same locality, 4.VI.1941, 1 3 ( MMBC); same locality, 23.VI.1942, 2 3 ( MMBC); same locality, 14.VIII.1942, 2 3 ( MMBC); Brno, Rudka, 18.V.1945, 1 3 ( MMBC); Horní Lomná, Hruška, 780m, 10–11.VIII.1997, M. Barták leg., 1 3 ( ZMUC); Hrubý Jeseník Mts., Velká Kotlina, 1200m, 7.VII.2004, M. Barták leg., 3 3 ( ZMUC); Kanice, 23.VII.1942, 2 3 ( MMBC); Lednice, 7.VI.1977, D. Povolný leg., 1 3 ( MMBC); Mnichovice, 350m, 20–21.VII.1996, M. Barták leg., 1 3 ( ZMUC); Pálava, 17.VII.2001, D. Povolný leg., 2 3 ( MMBC); St. Jan, 6.VII.1995, D. Povolný leg., 1 3 ( MMBC); Šumava, Malá Niva, 750m, 20–22.VII.1997, M. Barták leg., 1 3 ( ZMUC); Šumava, Popelná, 950m, 4.VII.1988, M. Barták leg., 1 3 ( ZMUC); Šumava-Plechý, 1050m, 22.VII.1992, M. Barták leg., 4 3 ( ZMUC); Vráž u Písku, 400m, 2.VI.1993, M. Barták leg., 1 3 ( ZMUC); same locality, 30.V.1994, M. Barták leg., 1 3 ( ZMUC); same locality, 1– 3.VI.2002, M. Barták leg., 1 3 ( ZMUC); Zaječí, 16–28.VIII.2002, D. Povolný leg., 5 3 ( MMBC). France: Hautes-Alpes, Montgenèvre, 1800m, 12.VII.1990, M. Barták leg., 1 3 ( ZMUC); Haute Savoie: nr. Ardent, la Lecherette, 22.VI.2000, T. Pape leg., 1 3 ( NHRS); Mont Ouson, above Col du Corbier, 27.VI.2000, T. Pape leg., 2 3 ( NHRS); Val-d’Oise, Carnelle, 10. VI.1911, 1 3 ( ZMUC). Germany: Weisbaden, 28.VI–3.VII.1909, 2 3 ( NHRS); same locality, 7.VI.1910, 1 3 ( NHRS). Hungary: Debrecen, R.T. Webber leg., 1 3 ( USNM); Kiskunság National Park, 23.VIII.1986, 1 3 ( ZMUC). Italy: Abruzzi, L’Aquila prov.: Anversa degli Abruzzi, Sorgente Cavuto, 520m, 30.VII.1997, P. Cerretti leg., 2 3 (CNBFVR); Barrea, Colle della Croce, 1200m, D. Whitmore et al. leg., 1 3 (CNBFVR); Opi, Valle Fredda, 1300m, beech forest, 22.V.2005, D. Whitmore et al. leg., 2 3 (CNBFVR); Chieti prov.: Abetina di Rosello, 940m, 21.V.2005, D. Whitmore et al. leg., 1 3 (CNBFVR); Palena, 1300m, 19.VIII.2000, M. Mei leg., 1 Ƥ ( MZUR); Emilia-Romagna, Parma prov., Corniglio, nr. Sella del Marmagna, 1747m, 10.VII.2010, D. Whitmore leg., 2 3 (CNBFVR); Latium, Rome prov.: Orvinio, La Spineta, 800m, 21.V.2000, M. Mei leg., 2 3 ( MZUR); Tenuta della Cervelletta, 4.V.1999, M. Mei leg., 1 3 ( MZUR); same locality, 19.IX.1999, M. Mei leg., 1 3 (CNBFVR); same locality, 22.IX.1999, M. Mei leg., 1 Ƥ ( MZUR); same locality, 9.X.1999, M. Mei leg., 1 3 ( MZUR); same locality, 2.IV.2000, M. Mei leg., 1 3 ( MZUR); same locality, 13.V.2000, M. Mei leg., 1 Ƥ ( MZUR); same locality, 3.VII.2000, M. Mei leg., 1 3 ( MZUR); Lombardy, Mantova, Marmirolo, Bosco della Fontana, 25m, 7.IX.2001, S. Vanin leg., 1 3 (CNBFVR); Trentino-Alto Adige, Trento prov., Storo, 9–16.VII.1986, S. Andersen leg., 1 3 ( ZMUC); Tuscany, Florence, 10–18.V.1986, T. Pape leg., 1 3 ( ZMUC). Norway: Kongsvoll, 12–20.VII.1985, O. Karsholt & V. Michelsen leg., 1 3 ( ZMUC). Russia: Leningrad region, several 3 ( USNM); Tyva, Eastern Sayan Ridge, Karzanak, nr. Mina, 6.VII.1959, Grunin leg., 1 3 ( YVC). Serbia: Deliblato National Park, 22.VII.1983, 2 3 ( ZMUC); Golubac, N. Baranov leg., several 3 ( USNM). Slovakia: Burda, 1–16.VII.1976, Čepelák leg., 3 3 ( MMBC); Hlohovec, 12.VIII.1989, Z. Mélek leg., 1 3 ( ZMUC). Spain: Costa Brava, Malgrat de Mar, 21.VI–4.VII.1996, D. Povolný leg., 16 3 ( MMBC); same locality [no date], D. Povolný leg., 6 3 ( MMBC); Lerida, Sorpe, 1200–1300m, 19–22.VI.1982, S. Andersen et al. leg., 1 3 ( ZMUC). Sweden: Abiskojokk, Ridonjira, 23.VII.1995, A.C. Pont leg., 1 3 ( NHRS). Switzerland: [no locality, no date] J.T. Skovgaard leg., 1 3 ( ZMUC); Glarus, Klöntal, Vorauen, 8.VIII.1995, B. Merz leg., 1 3 ( NHRS). Ukraine: Kosov, 28.V.1975, O. Victorova leg., 1 3 ( MMBC).

Remarks. The holotype of Hartigia anastrenua is conspecific with Sarcophaga (H.) vagans . I also agree with the recent synonymy of Sarcophaga bajkalensis with S. (H.) vagans proposed by Verves and Khrokalo (2006).

Figures

FIGURES 66 – 77. 66 – 68. Sarcophaga (Heteronychia) pseudobenaci (Baranov). 66. Cercus and surstylus, specimen from Greece, Hortiatis (CNBFVR), scale bar 0.15 mm. 67 – 68. Specimen from Greece, Stávros (ZMUC), scale bars 0.1 mm. 67. Distiphallus (lateral). 68. Juxta (dorsal). 69 – 71. Sarcophaga (Heteronychia) benaci Böttcher, specimens from Croatia, Krk (NHRS). 69. Cercus and surstylus, scale bar 0.1 mm. 70. Distiphallus (lateral), scale bar 0.05 mm. 71. Juxta (dorsal), scale bar 0.05 mm. 72 – 73. Sarcophaga (Heteronychia) infantilis Böttcher, specimen from Austria, Steinernes Meer (NHRS). 72. Cercus and surstylus, scale bar 0.1 mm. 73. Distiphallus (lateral), 0.05 mm. 74 – 75. Sarcophaga (Heteronychia) shnitnikovi (Rohdendorf), specimen from Kyrgyzstan, Kaindy (ZIN). 74. Cercus and surstylus, scale bar 0.25 mm. 75. Distiphallus (lateral), scale bar 0.1 mm. 76 – 77. Sarcophaga (Heteronychia) vagans Meigen, specimen from Austria, Niedere Tauern (ZMUC), scale bars 0.25 mm. 76. Phallus and gonites. 77. Cercus and surstylus.