Wormsina transsylvanica ( Hoernes & Auinger, 1880 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 49-50

Wormsina transsylvanica ( Hoernes & Auinger, 1880) nov. comb.

Figs 10F 1 –F 2 , G 1 –G 2 , H 1 –H 2 , I 1 –I 2 , J

Cylindra Transsylvanica nov. form.—Hoernes 1880: 125 [nomen nudum].

* Mitra (Cylindra) Transsylvanica nov. form.— Hoernes & Auinger 1880: 90, pl. 11, figs 1a–b, 2.

Mitra (Dibaphimitra) transsylvanica Hoernes & Auinger — Cernohorsky 1970: 70, pl. 3, fig. 8.

Mitra [(Dibaphimitra)] transsylvanica Hoernes & Auinger, 1880 — Cernohorsky 1976: 475, pl. 427, fig. 2.

Type material. Lectotype (designated herein): NHMW 1874 /0025/0008, SL: 27.8 mm, MD: 13.2 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1880, pl. 11, figs 1a–b), figs 10I 1 –I 2 . Paralectotypes: NHMW 2020 /0131/0001, SL: 26.3 mm , MD: 14.0 mm, Lăpugiu de Sus ( Romania) , illustrated in Hoernes & Auinger (1880, pl. 11, fig. 2), figs 10H 1 –H 2; NHMW 2020 /0132/0001, SL: 20.0 mm , MD: 11.1 mm, Lăpugiu de Sus ( Romania) , figs 10J; NHMW 2020 /0132/0002, SL: 29.6 mm , MD: 14.5 mm, Lăpugiu de Sus ( Romania) , figs 10F 1 –F 2; NHMW 2020 /0132/0003, SL: 28.5 mm , MD: 14.5 mm, Lăpugiu de Sus ( Romania) , figs 10G 1 –G 2; NHMW 1854 /0035/0227, 5 specimens, Lăpugiu de Sus ( Romania) .

Revised description. Shell medium sized, inflated ovoid with low, cyrtoconoid spire. Protoconch unknown. Teleoconch of five whorls; spire whorls broadly convex, with maximum diameter close to abapical suture; suture weakly incised. Last whorl high, ovoid, periphery slightly above mid-whorl, slowly contracting into short base. Early spire whorls covered by densely spaced spiral cords (about 15 on penultimate whorl) separated by narrow, deep grooves. Delicate, densely spaced axial ribs on first two teleoconch whorls forming cancellate, pitted pattern with spiral cords. Axial sculpture weakening rapidly abapically, absent by end of second whorl, sculpture abapically reduced to narrow pitted grooves, most prominent on adapical portion of last whorl. Periphery of last whorl nearly smooth; weak spiral cords separated by narrow pitted grooves re-appear on base and fasciole. Aperture moderately narrow, anteriorly widening, posteriorly narrowly angulated. Columella bearing four prominent oblique folds, weakening abapically; subobsolete fifth abapical fold present in some specimen. Columellar callus formed between adapical fold and siphonal canal, narrow, sharply delimited. Narrow chink between columellar callus and fasciole. Outer lip thin, without lirae or denticles. Siphonal canal short, wide, slightly recurved, with deep notch. Colour pattern in UV light consisting of spirally arranged vaguely subquadrate blotches separated by irregular, dark spiral bands on last whorl.

Shell measurements and ratios. SL = 20.0– 30.8 mm, MD: 11.1–14.6 mm, AA = 85–92°, SL/ MD: 1.9–2.0, AL/AW: 5.1–5.4, AH/S: 3.1–3.9.

Discussion. Mitra sp. aff. transylvanica [sic] Hoernes & Auinger 1880 described by Pfister & Wegmüller (2007) from the lower Miocene of Switzerland has blunt spiral cords and is clearly unrelated to Wormsina transsylvanica . Even its placement in Mitridae seems to be doubtful.

Palaeoenvironment. Unknown, the clays of the type locality Lăpugiu de Sus ( Romania) suggest offshore environments but the eroded surface and bioerosion of some specimens might also indicate transport from shallow water habitats .

Distribution in Central Paratethys. Badenian (middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania) ( Hoernes & Auinger, 1880).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 10. A1–A2. Fraudiziba mathiasi (Bałuk, 1997), Lăpugiu de Sus (Romania), NHMW 1854/0035/0085. B1–B2. Fraudiziba subcarinata (Bellardi, 1887), Modra-Kráľová (Slovakia), NHMW 2020/002/0001. C. Fraudiziba subcarinata (Bellardi,