Domiporta turpis, Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, The Mitridae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4983 (3), pp. 1-72 : 31-33

Domiporta turpis nov. sp.

Figs 7K 1 –K 2 , 8A 1 –A 2

Mitra scrobiculata Brocc. — Hörnes 1852b (pars): 100, pl. 10, fig. 17 [non Brocchi, 1814].

Type material. Holotype: NHMW 1846 /0037/0098a, SL: 24.5 mm, MD: 6.5 mm, Baden ( Austria), illustrated in Hörnes (1852b, pl. 10, fig. 17), 7K 1 –K 2, 8A 1 –A 2.

Type locality. Baden ( Austria), Vienna Basin .

Type stratum. Clay of the Baden Formation.

Age. Middle Miocene, middle Badenian (Langhian).

Etymology. From Latin turpis (= ugly).

Diagnosis. Domiporta species of medium size, slender fusiform profile, characterised by blunt, bifid ad- and abapical spiral cords on penultimate whorl and numerous, blunt, flattish, regularly spaced spiral cords on last whorl, delicate axial riblets in spiral grooves on spire and last whorl.

Description. Shell medium sized, slender fusiform with elongate last whorl. Protoconch unknown. Teleoconch of eight whorls. Spire whorls straight sided to weakly convex with deeply incised suture. First teleoconch whorls cancellate with two flattened spiral cords, increasing to three cords on fourth whorl. Ad- and abapical spiral cords bifid by intercalation of secondary spiral groove. Spiral grooves between primary cords deep, with rectangular cross section and delicate axial riblets, widening on penultimate and last whorls. Last whorl and base bearing about 18 blunt, broad, flattened spiral cords. Axial sculpture of delicate growth lines in deep interspaces. Last whorl slowly contracting with weak concavity at base. Aperture narrow, elongate. Columellar callus narrow, sharply delimited. Columella with two oblique, narrow, weak columellar folds; third abapical fold even weaker. Outer lip not preserved. Siphonal canal moderately long, narrow, slightly bent to left, weakly twisted, with deeply incised siphonal notch. Colour pattern in UV light ( Figs 7K 1 –K 2 ) consisting of dark spiral stripes coinciding with spiral grooves.

Shell measurements and ratios. SL = 22.5 mm, MD: 6.5 mm; AA = 30°, SL/MD: 3.7, AL/AW: 7.0, AH/S: 2.1.

Discussion. This species is characterised by its flattened spiral cords and deep spiral grooves. It is reminiscent of Domiporta austrogallica ( Mayer-Eymar, 1898) in shape and size, but differs in its more slender shape, coarser spiral sculpture, greater number of spiral cords on the last whorl, broader spiral cords, and the distinctly narrower siphonal canal. Domiporta sallomacensis sensu Peyrot, 1928 , from the Serravallian of France, seems to represent a closely related species, but is less slender, has more convex spire whorls, narrower spiral grooves, and lacks prominent axial riblets in the spiral grooves. Domiporta turpis nov. sp. could be mistaken for a juvenile Cancilla exornata (Bellardi, 1887) , but differs in its much smaller size and blunt spiral cords on early teleoconch whorls.

Palaeoenvironment. The clay of the Baden Formation formed in middle to outer neritic settings with up to 250 m water depth ( Hohenegger et al. 2008).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Baden ( Austria) ( Hörnes 1852b).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Palaeogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser & Landau 2017) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements for shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 7. Colour patterns in UV light. A1–A2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0002. B1–B2. Domiporta amoena nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0001. C1–C2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 1868/0001/0403. D1–D2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0003. E1–E2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0002. F1–F2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0003. G1–G2. Domiporta pulchra nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0001. H1–H2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0004. I1–I2. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus (Romania), NHMW 2020/0112/0001. J1–J2. Cancilla planicostata (Bellardi, 1887), Coşteiu de Sus (Romania), NHMW 2020/0112/0002. K1–K2. Domiporta turpis nov. sp., holotype, Baden (Austria), NHMW 1846/0037/0098ª. L1–L2. Episcomitra pseudoincognita nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0122/0001.

FIGURE 8. A1–A2. Domiporta turpis nov. sp., holotype, Baden (Austria), NHMW 1846/0037/0098a. B1–B2. Domiporta austrogallica (Mayer-Eymar, 1898), holotype, Baden (Austria), NHMW 1846/0037/0098. C1–C2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0002. D1–D2. Domiporta amoena nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0106/0001. E1–E2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania). F1–F2. Domiporta amoena nov. sp., paratype, Lăpugiu de Sus (Romania). G1–G2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0003. H1–H2. Domiporta pulchra nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0004. I1–I2. Domiporta pulchra nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 2020/0129/0001.