Ayacucho titschacki Roewer, 1949

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

publication ID	https://dx.doi.org/10.3897/asp.80.e73829
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scientific name	Ayacucho titschacki Roewer, 1949
status

Treatment

3.23. Ayacucho titschacki Roewer, 1949

Figs 3D View Figure 3 , 7U, V View Figure 7 , 12I, J View Figure 12 , 23E View Figure 23 , 28 View Figure 28

Ayacucho titschacki Roewer, 1949: 57 (desc.), figs. 114a (dorsal habitus), 114b (ventral habitus), 114c (chelicera, pedipalpus and carapace in lateral view); 114d (detail of coxa IV), 114e (sternum), 114f (chelicera in lateral view); Kury 2003: 144 (cat.).

Redescription.

MALE: Measurements (n =2) DSW: 3.0-4.1; DSL: 4.5-5.4; CL: 1.5-2.2. FIVL: 4.8-5.4. ChL: 2.5-4.0. Coloration: (Fig. 23E View Figure 23 ) Predominantly yellowish, with two longitudinal black stripes in the lateral region of dorsal scutum (carapace and lateral edges of the areas I-IV). Lateral margins of DS yellowish, with small black spots. Posterior margin of the dorsal scutum, free tergites and legs predominantly black. Chelicerae and pedipalpus predominantly dark yellow. Dorsum: (Fig. 3D View Figure 3 ) Alpha-type DSS. Anterior margin of carapace with median elevation totally covered with granules. Ocularium with two small acuminate tubercles, absent in some individuals. Areas I-IV with two pairs of small median tubercles; the lateral pair can be greatly reduced in some individuals. Posterior margin of dorsal scutum with a transverse row of 4-5 acuminate tubercles (larger than tubercles of areas I-IV). Free tergites I-III with a row of tubercles similar to the posterior margin of dorsal scutum, increasing the number of free tergite I to III. Chelicerae: (Fig. 3D View Figure 3 ) Swollen or not in males. Segment I densely covered with granules throughout its length. Segment II predominantly smooth, with some hairs on the front face; finger with three teeth. Segment III with two teeth. Pedipalpus : With very small granules sparsely distributed on the dorsal surface of the femur and patella. Trochanter with a distal ventral setiferous tubercle. Femur with a ventrobasal setiferous tubercle and a row of six small ventromedian setiferous tubercles. Tibia: prolateral IIII, retrolateral IIII. Tarsus: prolateral IIi, retrolateral Iii. Venter: Coxa I with dispersed small tubercles. Coxae II-III, predominantly smooth, with sparse granules. Coxa IV with granulation denser than coxae III-IV. Genital area smooth. Free sternites I-IV and anal operculum with small granules sparsely distributed. Legs: (Figs 3D View Figure 3 , 7U, V View Figure 7 ) Coxa I with a prolateral apophysis; coxa II with a retrolateral and a prolateral apophysis. Coxa IV densely granulate, with larger tubercles distributed more densely in the prolateral portion. Trochanters I-IV with few granules, unarmed. Femora I-III, with granules distributed throughout its length. Femur IV with granules densely distributed in the dorsal and lateral, less heavily on the ventral surface. Tarsal formula: (n =2) 7, 12, 7, 8. Penis: (Fig. 12I, J View Figure 12 ) VP rectangular; distal margin slightly concave, with short lateral-apical projections; slightly concave in lateral view. MS C1-C5 (or C6) subapical long and straight; MS A1 median short and straight; MS B1 sub basal long and straight (longer than MS A and shorter than MS C). Lateral sacs short and apically acuminated, with short T3-like microsetae. Stylus slightly thick, apically inflated, dorsally inclined, with apical tiny projections. Dorsal process absent. Promontory convex. - FEMALE: Measurements (n =4) DSW: 3.0-4.4; DSL: 4.5-5.4; CL: 1.9-4.7. FIVL: 3.7-4.5. ChL: 1.7-4.0. Swollen chelicerae in some specimens, like males. Tubercles of femur of pedipalpus slightly smaller. Ocularium tubercles smaller than in male. Granules on femur IV thinner when compared with those of male. Tarsal segmentation: (n =4) 6-7, 10-11, 7, 7-8.

Diagnosis.

Similar to Ayacucho glauberrochai sp. nov., A. pomacocha sp. nov., A. silvae sp. nov. and A. vargasllosai sp. nov. in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig. 3D View Figure 3 ). It differs from the previously mentioned species in the following combination of characteristics: ocularium with two small acuminate tubercles and densely granulate (Fig. 3D View Figure 3 ); male femur IV unarmed (Fig. 7U, V View Figure 7 ; unlike A. silvae sp. nov.); penis VP rectangular; five-six pairs of subapical MS C; one pair of MS A and B (Fig. 12I, J View Figure 12 ).

Remarks.

The male holotype and paratypes examined by Roewer (SMF RII 8589/20) are actually females.

Distribution.

(Fig. 28 View Figure 28 ) PERU. Ayacucho . Ayacucho , Ocollo and Virgem de Cacharras de Cocha.

Material examined.

Type material. Holotype ♀, ' PERU, Ayacucho , Ayacucho , without date and leg. (SMF RII 8589/20) - Paratype ♀, ‘ditto’ (SMF RII 8589/20). Additional material. 1 ♂, 2 ♀ ' PERU, Ayacucho , Virgem de Cacharras de Cocha , 13°01′47″S 73°52′14″W, without date and leg. (MZSP 36972) GoogleMaps . 2 ♀ ' PERU, Ayacucho , Near Ocollo , 13°19′53.0″S 74°30′17,6″ W, 29/IV/2011, R. Pinto-da-Rocha, A. Benedetti, J. Ochoa & D. Silva leg. (MUSM) GoogleMaps ; 1 ♂, 2 ♀, ‘ditto’ (MUBI); 1 ♂, 5 ♀ ‘ditto’ (MZSP 36973).