Anthracus mindanaoensis, Jaeger, 2017

Jaeger, Bernd, 2017, Five new species of the Anthracus annamensis group from the Philippines and India (Coleoptera, Carabidae, Harpalini, Stenolophina), Linzer biologische Beiträge 49 (1), pp. 585-608 : 590-594

publication ID	https://doi.org/ 10.5281/zenodo.5357351
persistent identifier	https://treatment.plazi.org/id/A71F87B6-E155-766E-FF5E-7454FD96FC0E
treatment provided by	Marcus
scientific name	Anthracus mindanaoensis
status	sp. nov.

Treatment

Anthracus mindanaoensis View in CoL nov.sp. ( Figs 8 View Figs 4-8 , 13, 14 View Figs 9-14 , 15-17 View Figs 15-17 )

T y p e m a t e r i a l: Holotype: 3 ( MCZ) labelled " Madaum R. Tagum / Mar 20-27 1931 / At trap lantern", " Davao Province / Mindanao, P. I. / C.F. Clagg coll", " C.E. White / Collection " and " HOLOTYPE 3 / Anthracus / mindanaoensis nov.sp. / des. B. Jaeger 2017" [red label].

The holotype is slightly teneral, with left antennomeres 7-11 and right antennomeres 8-11 missing.

Paratypes: 2 ♀♀ (MCZ) with same locality labels as the holotype, but one specimen additionally with "det. Darlington / at B.M. 1947-48 / Notes p.". All paratypes also labelled " PARATYPE ♀ / Anthracus / mindanaoensis nov.sp. / des. B. Jaeger 2017" [red label].

E t y m o l o g y: The species name refers to the distribution of the species.

D e s c r i p t i o n: General appearance as figured ( Fig. 15 View Figs 15-17 ). Body length 3.5-3.8 mm (HT 3.8 mm); width 1.3-1.4 mm.

Shiny, pronotum weakly, elytra moderately iridescent. Head, pronotum and elytra mainly dark brown (1 PT) or pale brown (HT + 1 PT), but clypeus paler reddish brown, labrum and mandibles (inner margins and apices of the latter blackish) pale reddish yellow or yellowish brown, pronotal lateral margins, hind angles and base, and elytral lateral margin and first interval pale reddish yellow or yellowish brown. Legs, palpi and antennomeres pale yellowish brown, or antennomeres 3-11 slightly infuscated (1 PT). Ventral surface mainly brown to yellowish brown, with head and proepisterna somewhat darker and epipleura somewhat paler.

Head ( Figs 15-17 View Figs 15-17 ) including eyes very broad (HW/PW: 0.78-0.80), with eyes moderately prominent (head 1.66-1.70 times as wide as head between eyes). Labrum rectilinear or slightly rounded at apical margin. Mandibles medium sized, not markedly prolonged and curved, left mandible moderately sharp at apex, not thickened or truncate. Antennae moderately long relative to pronotal length (AL/PL: 2.27-2.33) and rather short relative to elytral length (AL/EL: 0.78-0.80). Microsculpture on labrum almost isodiametric, on clypeus isodiametric to weakly transverse, on anterior half of head obliterated, or occa- sionally with traces of lightly to very lightly impressed isodiametric meshes, and on posterior half posterior to eyes with lightly impressed isodiametric meshes, becoming moderately transverse in front of pronotal anterior margin.

Pronotum ( Figs 15-17 View Figs 15-17 ) weakly transverse, 1.22-1.25 times as wide as long, 1.25-1.28 times as wide as head, widest in second quarter, lateral seta inserted about at beginning of second quarter. Apical margin almost straight or weakly emarginate. Anterior angles narrowly rounded at tips, weakly projecting forward. Sides weakly rounded to anterior angles, from widest point rectilinearly converging to posterior angles, or very faintly sinuate front of the latter, which are well developed and narrowly rounded at tips. Basal margin very slightly concave, almost rectilinear or weakly arcuate medially, and slightly oblique to posterior angles. Lateral furrows evenly narrow in apical half, becoming weakly widened in posterior third, where they are fused with the baso-lateral impressions. Baso-lateral impressions small to medium sized, clearly delimited from pronotal disc and the scarcely depressed median part of base, flattened to basal and lateral margin. Baso-lateral impressions and other parts of pronotal surface impunctate. Median line fine, disappearing before reaching basal and apical margins. Anterior transverse impression almost obliterated, or suggested. Microsculpture on disc with traces of very lightly impressed strongly transverse meshes, at baso-lateral impressions with weakly to moderately impressed isodiametric to moderately transverse meshes.

Elytra ( Fig. 15 View Figs 15-17 ) rather long,1.58-1.63 times as long as wide, 2.91-2.92 times as long and 1.44-1.48 times as wide as pronotum, with sides weakly widened posteriad, widest somewhat posterior to middle. Subapical sinuation weak, apical tip of each elytron rather sharp, in one specimen with a small dent. Elytral striae distinctly impressed and impunctate, scutellar striole long. Intervals rather flat to weakly convex on disc, becoming very slightly narrowed and moderately convex at apex. Basal pore at beginning of scutellar striole present, interval 3 in third quarter with one setiferous pore, adjoining stria 2. Microsculpture on scutellum isodiametric, on elytral intervals almost obsolete, only here and there traces of very lightly impressed transverse lines rudiments visible. Macropterous.

Metepisterna at inner margin about 1.6 times longer than wide at basal margin. Prosternum medially with 6, and in front of apical margin with a row of 6-8 longer setae. Prosternal process posteriorly with 1 long and distinct seta. Abdominal sternites IV-VI with distinct and rather dense pubescence, in sternite IV often with a small smooth area laterally. Last visible sternite of males faintly concave, and that of females almost rectilinear or very faintly convex at apex, at apical margin with two 2 setae in males and 4 in females.

Protarsomeres 1-4 of males moderately dilated and with distinct, biseriately arranged adhesive hairs on ventral surface. Mesotarsomeres 2-4 of males weakly dilated and with adhesive hairs on ventral surface. Pro- and mesotarsomere 4 of males not bilobed. In females pro- and mesotarsomeres unmodified. Male profemora not noticeable thickened in relation to females, and without numerous long and fine setae on upper inner margin.

Median lobe of aedeagus ( Figs 8 View Figs 4-8 , 13-14 View Figs 9-14 ) moderately large, with apical portion moderately long and gradually converging to the moderately broad tip ( Fig 13 View Figs 9-14 ). In lateral aspect only slightly curved ventrad with apical part rather straight ( Figs 8 View Figs 4-8 , 14 View Figs 9-14 ) and tip of apex with peculiar shape (8a). Internal sac (lateral aspect) composed of 1 large and 1 smaller subapical tooth, and in medial portion with one group of 2 larger and 6 smaller teeth arranged ventrally, and one group of 1 smaller, 1 larger, and 5 smaller teeth arranged dorsally.

N o t e s o n v a r i a t i o n: The holotype and one of the female paratypes agree quite well in external characters, particularly in the shape the pronotum ( Figs 15-16 View Figs 15-17 ). In contrast to that the second female has a slight sinuation in front of hind angles and their elytral tips are slightly denticulate. In these respects the specimen resembles on members of A. darlingtoni , although the presence of a small denticle at elytral apical tips varies also in this species from present to absent. However, whilst A. darlingtoni is so far only known from Leyte, this female was collected together with the holotype and the second female paratype, and it agrees with those in other characters, as small size and various body proportions. Therefore I include it in the type series of A. mindanaoensis , however, more material of both species is needed to confirm the boundaries of these two species.

C o m p a r i s o n s A. mindanaoensis belongs to the Anthracus annamensis group and represents the smallest of the three immaculate species which are now known from the Philippines. It is somewhat similar to the preceding species in general appearance (at least to the somewhat aberrant female paratype, see fig. 17) but it differs from A. darlingtoni by characters given under this species. From A. brevipennis nov.sp. it can be easily distinguished by the completely different habitus, but also by the extended pubescence of abdominal sternite IV (smooth, or with some single hairs in A. brevipennis ), the other chaetotaxy of male anal sternite, and the different size, shape and internal structures of the aedeagus.

From other immaculate Oriental species of the A. annamensis group which were revised some years ago ( JAEGER 2012), and occur in India, continental SE Asia and the Sunda islands, A. mindanaoensis differs by smaller size, and/or other habitus with rather narrow pronotum and elytra, the different shape and internal structures of aedeagus.

D i s t r i b u t i o n A. mindanaoensis is so far known only from the Philippine island of Mindanao. Because the species has fully developed hind wings it is probably more widely distributed, at least in the Philippines.