Immanus, Hulcr & Cognato, 2013

Lin, Wei, Li, You & Meng, Ling-Zeng, 2023, First record of the genus Immanus Hulcr & Cognato (Coleoptera: Curculionidae: Scolytinae: Xyleborini) from China, with description of a new species, Zootaxa 5352 (3), pp. 433-438 : 436-437

Key to species of the genus Immanus (females only)

Partly based on Beaver et al. (2019), Wang et al. (2020)

1 Elytral declivity with a distinct raised margin extending completely around the declivity............................. 2

- Elytral declivity with a costate but not raised margin extending partly or wholly around the declivity................... 7

2 Margin of elytral declivity with a series of spines or teeth extending from the upper margin to the apex................. 3

- Elytral declivity with raised margin crenulate or entire, without a series of distinct spines or teeth...................... 6

3 Striae and interstriae not distinguishable on declivity, declivital face with densely placed, confused granules, increasing in size from suture to margins of declivity, two or three small tubercles in mid-declivity in extension of discal interstriae 2. 5.1 mm long..................................................................... papuanus Beaver, Sittichaya & Liu

- Striae and interstriae distinct on declivity, declivital face without dense, confused granules. 5.0‒ 7.5 mm long............ 4

4 Elytral declivity matte, strongly impressed, concave above, lower half almost flat, margin with large pointed spines, largest in mid-declivity; declivital vestiture of short, bristle-like setae. Large species, 7.0‒ 7.5 mm long............ acanthurus (Lea)

- Elytral declivity shining, less strongly impressed, lower half weakly convex, marginal spines smaller and more equally sized; declivital vestiture of longer hairs. Smaller species, 5.0‒6.0 mm long............................................ 5

5 Elytral declivity umarmed on the upper half (ecxept marginal spines); declivital inerstriae 2 with three rows of setae on the upper half of declivity. Smaller species, 5.0 mm long....................................... duploarmatus (Browne)

- Elytral declivity armed with granules on the upper half (ecxept marginal spines); declivital interstriae 2 with two rows of setae on the upper half of declivity. Large species, 5.7‒6.0 mm long............................... pseudocolossus (Schedl)

6 Very large species, 8.5‒9.0 mm long. Declivity prolonged forwards so that its face is longer than wide, raised margin entire, not crenulate; interstriae 2 raised in middle of declivity with 3 or 4 tubercles on raised section…......... colossus (Blandford)

- Smaller species, 5.9‒6.0 mm long. Declivital face approximately circular, raised margin crenulate; interstriae 2 not distinctly raised, with a few small tubercles in middle of declivity..................................... permarginatus (Schedl)

7 Declivity abruptly steeply truncate, face circular, weakly convex, at least the sides acutely costate. Smaller species, 4.0‒ 4.5 mm long................................................................................................ 8

- Declivity strongly convex or obliquely truncate or steep and somewhat obliquely truncate, with a granulate costa extending to interstriae 7 only. Large species, 6.1‒7.8 mm long.............................................................9

8 Declivity opaque, covered with fine, setose granules on both striae and interstriae so that these are not easily distinguished; distinctly costate laterally, but not above; interstriae 2 not raised on declivity, with two larger tubercles in middle.................................................................................................. trolaki (Schedl)

- Declivity shining, with a finely granulate costa extending around the face of the declivity, confused granules present only near margins of declivity; declivital striae with large, closely placed, shallow punctures; interstriae 2 raised on declivity, with three widely spaced small pointed tubercles........................................................ desectus (Eggers)

9 Declivity rounded and strongly convex; two distinct large median tubercles on interstriae 2.......... sarawakensis (Eggers)

- Declivity steep and somewhat obliquely truncate or obliquely truncate; without large median tubercles on interstriae 2.... 10

10 Declivity obliquely truncate, interstriae bearing uniseriate small granules near summit, each granule bearing a long, erect hair-like seta. Large species, 7.8 mm long............................................. virago Wang, Smith & Cognato

- Declivity steep and somewhat obliquely truncate; interstriae bearing 2 or 3 rows of irregular uniformly granules along entire length, each granule bearing a long, recumbent hair-like seta. Smaller species, 6.1 mm long................ songi sp. nov