Quedius, Stephens, 1829, Stephens, 1829

Smetana, Aleš, 2012, Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 42. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 10, Zootaxa 3156, pp. 43-68 : 66

publication ID	https://doi.org/ 10.11646/zootaxa.3156.1.2
DOI	https://doi.org/10.5281/zenodo.5910944
persistent identifier	https://treatment.plazi.org/id/A43A6804-FFA6-C706-4BC5-4F07FA4EB39D
treatment provided by	Plazi
scientific name	Quedius
status

Treatment

Key to winged species of the muscicola- group View in CoL of mainland China

1. Legs uniformly testaceous............................................................................... 2

- At least tibiae of hind legs distinctly darkened, sometimes tibiae of middle legs and femora and tibiae of hind legs distinctly darkened............................................................................................ 5

2. Aedoeagus and paramere of characteristic shape ( Figs. 25, 26 View FIGURES 23 – 31. 23 – 26 ), sensory peg setae on underside of paramere quite numerous, situated as in Fig. 26 View FIGURES 23 – 31. 23 – 26 . Visible abdominal tergites one through five each with impression at each side laterad of midline, impressions becoming gradually less distinct toward apex of abdomen. Punctation of abdominal tergites in general moderately dense (details in description). Length 5.5 mm ..................................................... Q. hegesias View in CoL sp. nov.

- Aedoeagi and parameres of different shapes, with differently situated sensory peg setae. Impressions on abdominal tergites, if present, inconspicuous, mostly best observable on first three visible tergites. Punctation of abdominal tergites in general sparse to very sparse (details in description)...................................................................... 3

3. Apical portion of median lobe of aedoeagus narrow, with apex subacute ( Figs. 12, 13 View FIGURES 11 – 22. 11 – 15 ). Length 4.8 mm ... Q. filiolus View in CoL sp. nov.

- Apical portion of median lobe of aedoeagus variably wide, with apex arcuate to variably rounded ( Figs. 4–6 View FIGURES 1 – 10. 1 – 9 )............ 4

4. Aedoeagus pale, paramere narrow, usually more or less narrowed anteriad ( Figs. 4, 7, 8 View FIGURES 1 – 10. 1 – 9 ). Sensory peg setae on underside of paramere forming two lateral longitudinal rows ( Figs. 7, 8 View FIGURES 1 – 10. 1 – 9 ). Pronotum about as long as wide, no more than moderately narrowed anteriad. Punctation of elytra fine and dense. Length 4.8–6.0 mm........................ Q. muscicola Cameron View in CoL

-... Aedoeagus dark, paramere relatively wide, largely parallelsided. Sensory peg setae on underside of paramere forming one medial longitudinal group ( Fig. 21 View FIGURES 11 – 22. 11 – 15 ). Pronotum narrow, markedly narrowed anteriad. Punctation of elytra coarse and sparse. Length 4.8 mm .......................................................................... Q. hecato View in CoL sp. nov

5. First three visible abdominal tergites each with three shallow, inconspicuous impressions at base (middle impression sometimes missing); each lateral impression with slightly denser punctation and pubescence than on rest of surface............ 6

- First two visible abdominal tergites each with inconspicuous impression at each side, or tergites missing any impressions... 7

6. Median lobe of aedoeagus anteriorly dilated before narrowed into apical portion ( Figs. 45, 47 View FIGURES 43 – 54. 43 – 50 ). Paramere narrow, parallelsided ( Fig. 45 View FIGURES 43 – 54. 43 – 50 ). First four segments of male front tarsus markedly dilated, second segment as wide as apex of tibia. Length 4.3–5.0 mm........................................................................... Q. impressiventris View in CoL sp. nov.

- Median lobe of aedoeagus anteriorly not dilated before narrowed into apical portion ( Figs. 57, 58 View FIGURES 55 – 65. 55 – 61 ). Paramere wider, fusiform ( Figs. 56, 59 View FIGURES 55 – 65. 55 – 61 ). First four segments of male front tarsus slightly dilated, second segment narrower that apex of tibia. Length 4.7– 5.0 mm......................................................................... Q. tergimpressus View in CoL sp. nov.

7. Pubescence of elytra golden-yellowish.................................................................... 8

- Pubescence of elytra dark, brownish to dark brownish....................................................... 10

8. Middle and hind tibiae and hind femora darkened. First four segments of male front tarsus hardly dilated, segment 2 narrower than apex of tibia (ratio 0.85). Aedoeagus as in Figs. 80, 81. Length 3.9–4.2 mm ...................... Q. dryas View in CoL sp. nov.

- Only hind tibiae darkened. First four segments of male front tarsus markedly dilated, segment 2 slightly wider than apex of tibia (ratios 1.14 and 1.15). Aedoeagi different ( Figs. 65 View FIGURES 55 – 65. 55 – 61 , 72 View FIGURES 66 – 76. 66 – 68 )................................................... 9

9. Median lobe of aedoeagus very narrow, in almost straight line narrowed into extremely narrow apical portion with quite acute apex ( Figs. 65 View FIGURES 55 – 65. 55 – 61 , 66 View FIGURES 66 – 76. 66 – 68 ). Coloration of pronotum sexually dimorphic, dark brown becoming paler toward lateral margins in male, black in female. Length 4.0– 4.8 mm ........................................................ Q. meilixue View in CoL sp. nov.

- Median lobe of aedoeagus moderately narrow, parallelsided, anteriorly narrowed into short apical portion with acute apex ( Figs. 72, 73 View FIGURES 66 – 76. 66 – 68 ). Coloration of pronotum black in both sexes. Length 4.0– 4.5 mm ........................ Q. zetes View in CoL sp. nov.

10. Aedoeagus long, very narrow; paramere quite narrow, elongate, more or less parallelsided to narrowly fusiform, with narrowly arcuate apex not quite reaching, or reaching, apex of median lobe ( Figs. 31 View FIGURES 23 – 31. 23 – 26 , 34, 35 View FIGURES 32 – 42. 32 – 36 ). Tergite 10 of male genital segment shaped as in Fig. 28 View FIGURES 23 – 31. 23 – 26 , sparingly setose in front of long setae at apical margin. Sternite 9 of male genital segment with general setae numerous, short ( Fig. 29 View FIGURES 23 – 31. 23 – 26 ). Length 4.8–5,4 mm. Widely distributed species................... Q. maculiventris Bernhauer

- Aedoeagus markedly shorter, less narrow; paramere wider, fusiform, with narrowly arcuate apex not reaching apex of median lobe ( Figs. 40, 41 View FIGURES 32 – 42. 32 – 36 ). Tergite 10 of male genital segment shaped as in Fig. 38 View FIGURES 32 – 42. 32 – 36 , with only two short setae on middle portion in front of long setae ( Fig. 38 View FIGURES 32 – 42. 32 – 36 ). Sternite 9 of male genital segment with general setae sparse, markedly long ( Fig. 39 View FIGURES 32 – 42. 32 – 36 ). Length 4.9 mm. Eastern Sichuan: Huaying Shan..................................................... Q. optabilis Bernhauer View in CoL