Allodia dibolia Magnussen

Magnussen, Trude, Soli, Geir E. E. & Kjaerandsen, Jostein, 2019, Allodia Winnertz from the Himalayas, with nine species new to science (Diptera, Mycetophilidae), ZooKeys 820, pp. 119-138 : 123-125

Allodia dibolia Magnussen sp. n. Fig. 2B

Diagnosis.

The dorsal lobe of the gonostylus is tapering towards apex, and the median lobe is distinctly flattened and broad. Additionally, the basal lobe of the gonostylus is large, flat with many thick, short setae. All studied specimens of A. dibolia are small (<2.7 mm), compared to other species described from the Himalayan range. The specimens are all fairly uniform in their colouration patterns, but some are darker than others.

Type locality.

NEPAL: Province no. 1 (Kosi Zone), Terhathum District, Basantapur, 2300 m a.s.l.

Type specimens.

Holotype: male. 3 printed labels: (E. NEPAL) Basantapur (2300 m) 27°06'N, 87°23'E --- 27°08'N, 87°26'E / May 6, 1972 J. Emoto leg. Kyushu Univ. Col. / TSZD-JKJ-104944 (KUEC; pinned with genitalia in separate microvial). Paratypes: NEPAL: Province no. 1 (Kosi Zone), Terhathum District, Basantapur, 27°06'N, 087°23'E, 2300 m a.s.l, 29 Apr 1972, Leg. H. Shima, TSZD-JKJ-104946, male (KUEC), TSZD-JKJ-104947, male (TMU); 9 May 1972, Leg. H. Shima, TSZD-JKJ-104948, male (KUEC); NEPAL: Bhojpur District, Chiaksila, 27°26'N, 086°57'E, 2730 m a.s.l, 16 Jul 1981, Leg. J. Emoto, TSZD-JKJ-104949, male (KUEC); NEPAL: Sankhuwasabha District, Salpa La, 27°27'N, 086°55'E, 3000-3050 m a.s.l, 29 Jul 1981, Leg. J. Emoto, TSZD-JKJ-104950, male (KUEC); NEPAL: Province no. 3 (Nepalmandal, Bagmati), Nuwakot District, Sheopuri Lekh (= Siwapuri), Shivpuri Nagarjun NP, 27°48'N, 085°22'E, 2460 m a.s.l, 28 May 1981, Leg. J. Emoto, TSZD-JKJ-104951, male (KUEC); NEPAL: Province no. 3 (Sagarmatha Zone), Solukhumbu District, Junbesi Khola, 27°36'N, 086°33'E, 3400-3500 m a.s.l, 12 Aug 1981, Leg. J. Emoto, TSZD-JKJ-104952, male (KUEC); BHUTAN: Paro District, Jilay La, 27°22'11"N, 089°20'47"E, 3800 m a.s.l, 19 Aug 1993, Leg. T. Saigusa, TSZD-JKJ-104945, male (TMU).

Description. Body length 2.2-2.7 mm; wing length 2.2-2.8 mm. Colouration. Head and clypeus dark brown. Mouthparts, including palpomeres yellow. Antennae brown, with scape, pedicel, and base of first flagellomere yellow. Scutum brown, with narrow yellow lateral area, from humerus towards wing base. Antepronotum yellow, other lateral sclerites brown. Wings clear without markings. Halteres yellow. Legs yellow. Abdomen brown, tergites II-IV with whitish yellow area, larger towards posterior margin. Terminalia yellow. Head. Three ocelli present, median ocellus smaller than laterals, lateral ocelli touching eye margin. Head covered with fine trichia, except for few setae near eye margin. Antennae approx. twice as long as thorax. Scape and pedicel with several setae on distal third. Flagellomeres cylindrical, densely clothed with fine trichia. First flagellomere almost twice as long as pedicel. Thorax. Antepronotum with four strong setae. Scutum covered with uniform small, pale setae; strong prealar and postalar setae. Discal bristles absent. Scutellum with two strong bristles. Laterotergite with three short setae. Other lateral sclerites bare. Legs. All tibiae with short setae ar ranged in rows. Mid tibia with four long anterodorsal and ca. 20 short posterodorsal bristles. Hind tibia with eight anterodorsal and five posterodorsal bristles. Wings. Sc short, ending in R. Length of rm equal to stem of posterior fork. Base of anterior fork well before base of posterior fork. R1 with setae, R5 without setae. Male terminalia. Tergite IX medially divided, each part rounded covered with minute trichia, with one strong apical bristle. Hypandrial lobe heavily sclerotised, elongated and pointed inwards. Gonostylus with dorsal lobe apically tapering; outer surface with several strong setae, in particular near basis. Median lobe flat and broad, larger than dorsal lobe, but more weakly sclerotised; with mixture of short and long setae along posterior edge, some setae on inner surface of lobe apically. Ventral lobe short, club-shaped; with three strong setae apically. Basal part large, with straight edge, lined with strong setae. Internal part with long, caudally projecting process.

Etymology.

From Greek, dibolia, a two-edged lance, referring to the shape of the dorsal lobe of the gonostylus, which is spear-like.

Remarks.

Allodia dibolia is very distinct, and the gonostylus clearly separates it from other species of the genus. Superficially the terminalia can resemble those of A. lugens (Wiedemann, 1817), with a distinctly pointed dorsal lobe of the gonostylus. Allodia dibolia , however, the inner lobe of the gonostylus possess a projecting process, which is lacking in A. lugens . The medial lobe is large and broad, not pointed, with many setae, not only along the edge.