Albunea symnista (Linnaeus, 1767)

BOYKO, CHRISTOPHER B., 2002, A Worldwide Revision Of The Recent And Fossil Sand Crabs Of The Albuneidae Stimpson And Blepharipodidae, New Family (Crustacea: Decapoda: Anomura: Hippoidea), Bulletin of the American Museum of Natural History 2002 (272), pp. 1-396 : 260-296

Albunea symnista [sic]: Gordon, 1938: 187 (part)*

(not Albunea symmysta ( Linnaeus, 1758)) .? Albuneidae sp. DiSalvo et al., 1988: 458.

MATERIAL EXAMINED: Taiwan: Ma­Kung, Peng­Hui Island, Sept. 15, 1996, coll. unknown: 1 ♀, 17.4 mm cl, paratype ( NTOU).

Australia: Queensland: Samovrez Reef , 8 m, Sept. 1974, coll. N. Coleman: 1 ♀, 19.3 mm cl, paratype (AM P20456) ; Cape Bowling Green , 16 fms (= 29.3 m), Nov. 23, 1962, coll. W. Goode on R/V ‘‘ Dorothea’ ’: 1 Ƌ, 11.8 mm cl, 1 ♀, 17.9 mm cl, paratypes ( WAM 23387) ; New South Wales: Off Richmond River mouth, off Ballina , 28°52̍S, 153°34̍E, 16 m, Oct. 6, 1962, coll. unknown: 1 Ƌ, 15.4 mm cl, holotype (AM P15353) ; Lord Howe Island , 31°33̍S, 159°05̍E, coll. unknown: 1 ♀, 18.9 mm cl, allotype (AM P1925) .

New Caledonia: ‘‘New Caledonia,’’ 1903, coll. A. Milne Edwards: 1 ♀, 19.4 mm cl (MNHN­Hi 150).

Fiji: Viti Island: 1 Ƌ, 7.5 mm cl, paratype ( ZMH K–5137 ex Museum Goddefroy 2120).

Pitcairn Island: Sta. PIT­UI, Haul 13, 1 mi northwest of Pitcairn Island, 25°02–04̍S, 130°06–07̍W, 26–30 fms (= 47.6–54.9 m), Sept. 20, 1967, coll. National Geographic Society–Smithsonian­Bishop Museum Marquesas Expedition: 1 unsexable anterior half with pereopod I, 17.7 mm cl ( USNM 304304).

DIAGNOSIS: Carapace wider than long, covered with strongly setose grooves. Anterior margin with 8–10 spines on either side of ocular sinus. Setal field with thin lateral elements and concave anterior margin. CG1 with separate posterior lateral elements; CG4 with three or four short medial elements between longer supralateral elements of CG4, CG5 two short, irregularly convex, transverse elements; CG6 and CG7 separate; CG8 present; CG11 present. Rostrum present, reaching posterior margin of ocular plate. Ocular plate subquadrate. Distal peduncular segments dorsoventrally flattened and oblong in shape, rounded at tip, approximated along mesial margins, lateral margins convex, mesial margins straight. Cornea on lateral margin at tip. Dactylus of pereopod II with heel produced and rounded. Dactylus of pereopod III with heel produced and rounded. Dactylus of pereopod IV evenly sinuous from base to tip, with shallow indent. Telson of male divided into buttonlike proximal two­thirds and mucronate, tapering, distal third; proximal two­thirds rounded, heavily calcified, dorsally inflated, row of long setae flanking distal half of telson, medial transverse row of long setae present; distal third of telson weakly calcified, sharply tapering distally with produced tip. Telson of female ovate, tapering slightly distally, tip slightly produced.

DESCRIPTION: Carapace (fig. 85A) wider than long. Anterior margin faintly concave on either side of ocular sinus, becoming convex laterally with 8–10 large spines (n = 5) on each side along length. Rostrum a small acute tooth, reaching proximal margin of ocular plate. Ocular sinus smoothly concave. Frontal region smooth; setal field broad anteriorly and narrow posteriorly; posterior lateral elements reduced to narrow bands of setae. CG1 parallel to anterior margin of carapace, convex, sinuous, strongly crenulate, divided into medial fragment and curved, posteriorly displaced lateral elements. Mesogastric region smooth; CG2 present as two or three short elements, anteriorly and posteriorly displaced; CG3 broken into three to six short curved elements between posterior lateral elements of CG1, medial two elements shortest; CG4 with three or four short medial elements between longer supralateral elements of CG4, medial one or two elements anteriorly displaced, lateral short elements posteriorly displaced. Hepatic region smooth, with oblique setose groove at median of lateral margin, nearly reaching posterior element of CG1. Epibranchial region generally triangular, smooth; posterolateral margin with one or two oblique rows of short setae. Metagastric region smooth; CG5 present as two short, irregularly convex, transverse elements. CG6 strongly crenulate, strongly anteriorly concave medially, and sloping out to anteriorly convex lateral thirds. CG7 oblique, not reaching lateral margins of median segment of CG6. Cardiac region smooth; CG8 present as one long medial element. CG9 present as two short lateral grooves with broad gap at midline. CG10 present as two short lateral elements. CG11 present as two or three short elements. Post­ CG11 element absent. Branchial region with numerous short and long transverse rows of setae in anterior half. Posterior margin deeply and evenly convex, with submarginal groove reaching one­third up margin of posterior concavity. Branchiostegite with strong anterior submarginal spine; anterior region with scattered short, transverse lines ventral to linea anomurica; with many short rows of setae and sparsely covered with long plumose setae ventrally; posterior region membranous, with numerous irregular fragments and sparsely covered with long plumose setae.

Ocular plate (fig. 85B) triangular with deep median indentation; median peduncular segments present as small ovate, calcified areas lateral to ocular plate. Distal peduncular segments elongate, subtriangular, with convex lateral and straight mesial margins, cornea covering lateral portion of distal tip; lateral margins with faint notch one­fourth distal from base; mesial margins approximated along length; lateral and mesial margins with long plumose setae; tuft of plumose setae at proximolateral ventral angles and ventromedial longitudinal row of plumose setae extending from tuft to three­fourths length of peduncle.

Antennule (fig. 85C) with segment III narrow proximally, expanding distally to three times proximal width; plumose setae on dorsal and ventral margins and sparsely scattered on lateral surface; dorsal exopodal flagellum with 67–77 articles (n = 2), long plumose setae on dorsal and ventral margins; ventral endopodal flagellum with three or four articles (n = 4), plumose setae on dorsal and ventral margins. Segment II medially in­ flated in dorsal view, with plumose setae on dorsal and ventral margins and scattered on mediolateral surface. Segment I wider than long, unarmed; dorsal quarter of lateral surface rugose, with long plumose setae; long plumose setae on dorsal and ventral margins.

Antenna (fig. 85D) with segment V approximately 2.5 times longer than wide, with long plumose setae on dorsal and distal margins and scattered on lateral surface; flagellum with six articles (n = 5), long plumose setae on dorsal, ventral, and distal margins. Segment IV expanded distally, with long plumose setae on dorsal, ventral, and distal margins, and interrupted row of short setae on dorsolateral surface. Segment III with long plumose setae on ventral margin; short simple setae on dorsal margin and scattered on surface. Segment II short, widening distally, dorsally rugose, with plumose setae on margins and short simple setae scattered on lateral surface; antennal acicle long, thin, and reaching distal margin of segment IV, with long plumose setae on dorsal margin. Segment I rounded proximally, flattened ventrolaterally, with long plumose setae on dorsal and distoventral margins, and short simple setae on surface rugae behind spine; lateral surface with acute spine dorsodistally; low semicircular dorsolateral lobe ventrodistal to spine, margin of lobe with long plumose setae; segment with ventromesial antennal gland pore.

Mandible (fig. 85E) incisor process with two teeth; cutting edge with one tooth. Palp three­segmented, with plumose setae on margins and long, thick, simple setae arising from bend in second segment and on distal margin of terminal segment.

Maxillule (fig. 85F) distal endite proximally narrow, widening to inflated distal end, with thick simple setae on distal margin and thin simple setae on dorsal margin. Proximal endite with thick simple setae on distal margin. Endopodal external lobe truncate distally and curled under; internal lobe reduced with five thick setae at distolateral margin and one thick seta in median of lobe.

Maxilla (fig. 85G) exopod evenly rounded, with plumose setae along distal margin. Scaphognathite bluntly angled on posterior lobe, with plumose setae.

Maxilliped I (fig. 85H) epipod with plu­ mose setae on margins, distolateral surface, and mesial surface. Endite tapered distally and subequal to first segment of exopod. Exopod with two segments: proximal segment narrow, margins parallel, with long plumose setae; distal segment spatulate, longer than wide, broadest medially, margins and mesiodorsal surface with long plumose setae. Endopod flattened and elongate, reaching twothirds of distance to distal end of proximal exopodal segment; plumose setae on margins and median of lateral surface.

Maxilliped II (fig. 85I) dactylus evenly rounded, length subequal to width, with thick simple setae distally and on distolateral surface. Propodus two times wider than long, slightly produced at dorsodistal angle, with plumose setae on dorsal margin and patch of long simple setae on dorsodistal and ventrodistal angles. Carpus not produced dorsodistally, approximately two times longer than wide; long simple setae on dorsal margin and in patches on dorsodistal and ventrodistal angles. Merus approximately three times longer than wide, margins parallel; with long simple and plumose setae on margins and scattered on lateral surface. Basis­ischium incompletely fused, with plumose setae on margins. Exopod 1.7 times longer than merus, flagellum with one elongate article, approximately as long as carpus.

Maxilliped III (fig. 85J) dactylus oblong with rounded tip; long plumose setae on margins and in medial row on lateral surface. Propodus dorsodistally inflated, with longitudinal median row of plumose setae on lateral surface and three or four short patches of long plumose setae submarginally ventrally; dorsal margin with long plumose setae. Carpus produced onto propodus approximately two­thirds length of propodus; lateral surface with two transverse rows of long plumose setae; long plumose setae on dorsal and distal margins. Merus inflated, unarmed, with long plumose setae on margins and few scattered small patches on lateral surface. Basisischium incompletely fused, with weak crista dentata of four teeth. Exopod two­segment­ ed: proximal segment small; distal segment styliform, bluntly rounded distally, approximately one­third length of merus; with plumose setae on margins; without flagellum.

Pereopod I (fig. 86A) dactylus curved and tapering; lateral and mesial surfaces smooth; dorsal margin with long plumose setae; ventral margin with short simple setae. Propodal lateral surface with numerous short, transverse rows of setose rugae; dorsal margin unarmed; ventral margin distally produced into acute spine; cutting edge lacking teeth, lined with long plumose setae; dorsal margin with long plumose setae, ventral margin with short simple setae. Carpus with dorsodistal angle produced into rounded projection, small dorsally directed acute spine on distodorsal mesial margin; dorsal margin with short transverse grooves behind distal projection; dorsal and distal margins with short plumose setae; lateral surface with small distal rugose area, with few transverse setose ridges on distal half of surface; mesial surface smooth with medial transverse row of setae, margins with long plumose setae. Merus unarmed; lateral surface with scattered transverse rows of long plumose setae, distal and mediodorsal margins with long plumose setae; mesial side with few scattered setae; fully calcified. Basis­ischium incompletely fused, unarmed. Coxa unarmed.

Pereopod II (fig. 86B) dactylus smooth; base to heel faintly convex, heel low and rounded, subquadrate, heel to tip with broad, subacute indent, tip subacute, tip to base broadly convex; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, several widely spaced submarginal tufts of short setae dorsodistally; mesial surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae, with patch of long plumose setae at base. Propodal dorsal surface smooth, with ventral margin inflated and rounded; oblique row of long plumose setae on distal margin of lateral surface; distal and ventral margins with long plumose setae; dorsolateral surface as narrow, oblique, flattened shelf, with short setae on dorsal margin and long plumose setae on ventral margin; mesial surface with elevated, curved setose ridge from ventral junction with dactylus almost to ventral proximal junction with carpus. Carpus strongly produced, tapered distally, and rounded dorsodistally, dorsal margin smooth; lateral surface smooth, produced area smooth, irregular, interrupted row of rugae and submarginal elevated ridge ventrally, rugae and ridge with long plumose setae; dorsal margin with short plumose setae, ventral margin with long plumose setae; mesial surface smooth, with row of long plumose setae distally, ventrally, and subdorsally. Merus with large median decalcified window covering nearly all of lateral surface, long plumose setae on dorsodistal and ventral margins, few scattered long plumose setae on surface; mesial surface nearly smooth with two long rows of setae. Basisischium incompletely fused and unarmed. Coxa unarmed.

Pereopod III (fig. 86C) dactylus with base to heel straight, heel low and subquadrate, heel to tip with broadly concave indent, tip subacute, tip to base smoothly convex; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, dorsodistal margin with tufts of short setae; ventral margin with long plumose setae, dorsal margin with short simple and plumose setae; mesial surface smooth, with plumose setae proximally at junction with propodus. Propodus not inflat­ ed dorsoventrally; lateral surface smooth, with long plumose setae in oblique row, simple setae on dorsal margin, plumose setae on ventral margin; dorsolateral surface narrow, oblique, flattened, without setose mat; mesial surface smooth. Carpus produced dorsodistally, exceeding proximal margin of propodus by one­half length of propodus; tip subacute, dorsal margin unarmed; lateral surface slightly rugose dorsodistally, with mat of short setae and two interrupted rows of setae ventrally; mesial surface smooth, with long plumose setae on distal margin and in short oblique row on distal surface. Merus smooth, narrow, and almost cylindrical, with large decalcified window covering nearly half of lateral surface medially; dorsal and ventral margins unarmed, dorsodistal and ventral margins with long plumose setae; mesial surface smooth. Basis­ischium incompletely fused and unarmed. Coxa unarmed. Female with large gonopore on anterior mesial margin of coxa, surrounded with short plumose setae; male with small pore.

Pereopod IV (fig. 86D) dactylus with base to tip convex proximally, with indistinct heel and indent and almost straight from indent to tip, tip acute, tip to base concave distally to convex proximally; lateral surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae; mesial surface with dorsal decalcified region, demarcated ventrally by longitudinal elevated ridge with row of short setae; with setose punctations ventral to decalcified window. Propodus expanded dorsally and ventrally; ventral expansion exceeding ventral margin of dactylus, ventral margin with long plumose setae; dorsal expansion with row of long plumose setae dorsally, oblique area with mat of short simple setae; lateral surface smooth, mesial surface smooth, with distoventral area of few patches of long plumose setae. Carpus slightly produced dorsodistally; ventral four­fifths of lateral surface and mesial surface smooth, dorsodistal one­fifth of lateral surface with mat of short simple setae; dorsal margin with short simple and long plumose setae; ventral margin with short simple setae; mesial surface decalcified medially. Merus lateral surface with scattered short transverse rows of setae, dorsal margin with long plumose setae; proximoventral fourth of mesial surface with large decalcified window. Basis­ischium incompletely fused and unarmed. Coxa unarmed.

Abdomen (fig. 86E) somite I wider than long, widest posteriorly; dorsal surface with anterior margin straight; posterior margin curved, with elevated submarginal row of short setae; small transverse decalcified windows laterad of segment median. Somite II dorsal surface with irregular, submarginal, transverse ridge anteriorly; with small, transverse, decalcified windows laterad of segment median just anterior to submarginal ridge; pleura expanded and directed laterally; anterolateral margins rounded, anterior and lateral margins with long plumose setae, posterolateral angle rounded, posterior margin with short setae; posteromesial angle with mat of short simple setae. Somite III similar to somite II, but narrower, shorter, anterior submarginal windows present; pleura thinner and shorter than on somite II, directed posterolaterally proximally and anterolaterally distally, with setae as in somite II; anterolateral angle subacute; dorsal surface obliquely flattened anterolaterally, with submarginal row of short setae. Somite IV similar to somite III, but thinner and shorter, anterior submarginal windows present; pleura thinner and shorter than on somite III, directed laterally; dorsal surface obliquely flattened anterolaterally; lateral and posterior margins with long plumose setae, anterior margin with short simple setae. Somite V wider than somite IV, anterior submarginal windows present; lateral margins with plumose setae; pleura absent. Somite VI slightly broader than somite V, anterior submarginal windows present; dorsal surface with two short transverse rows of setae laterad of midline and on posterior margin; pleura absent.

Females with uniramous, paired pleopods on somites II–V; males without pleopods.

Telson of male (fig. 86F) divided into a subcircular (‘‘buttonlike’’) proximal twothirds and mucronate tapering distal third; proximal two­thirds rounded, heavily calcified, dorsally inflated, with deep median groove and long plumose setae flanking distal third of groove to distal end of calcified region, proximolateral angles with few short simple setae, transverse row of long plumose setae present on either side of median groove approximately one­third proximal from distal margin of calcified region; distal third of telson weakly calcified, sharply tapering distally with produced tip; entire telson lined with long plumose setae on margins in distal twothirds. Telson of female (fig. 86G) ovate, tapering slightly distally, tip slightly produced, dorsal surface evenly calcified; median groove present in medial two­thirds of dorsal surface, lined with short simple setae; proximolateral angle with few short simple setae, margins with long simple setae.

DISTRIBUTION: Known from Taiwan; Queensland, Australia; New Caledonia; and Fiji, in up to 29.3 m depth. Also provisionally from Pitcairn Island, in 47.6–54.9 m depth.

MAXIMUM SIZE: Males: 15.4 mm cl; females, 19.4 mm cl.

TYPE SPECIMENS: AM P15353 (holotype), AM P1925 (allotype), AM P20456 (paratype), NTOU (paratype), WAM 23387 (2 paratypes), ZMH K–5137 (paratype).

TYPE LOCALITY: Off Richmond River mouth, off Ballina, New South Wales, Australia, 28°52̍S, 153°34̍E, 16 m.

ETYMOLOGY: The specific name of this taxon is given for the distinctive buttonlike morphology of the telson of the males.

REMARKS: The Pitcairn specimen exists only as the carapace and anterior half of the animal with all appendages anterior to and including pereopod I. Given its poor condition, it is only provisionally referred to this species, although the characters of the extant structures are essentially identical. The single reference to an Easter Island albuneid (Di­ Salvo et al., 1988) may also be this taxon, as there are several crustacean species which are known from both Easter Island and Pitcairn Island (Boyko, in prep.). Despite repeated attempts to collect albuneids on Easter Island over a 3–week period in August 1999, I found no specimens.

This species belongs to the ‘‘ carabus ­ group’’ of Albunea and is the sister taxon to the other two species in that group. The distinctive ‘‘buttonlike’’ morphology of the male telson separates it from both A. carabus and A. danai .

Albunea carabus ( Linnaeus, 1758) Figures 87 , 88

Cancer Carabus Linnaeus, 1758: 632 . – Linnaeus, 1767: 1052. – Houttuyn, 1769: 417. – de Villers, 1789: 156.

Cancer carabus: Statius Müller, 1775: 1126– 1127 . – Latreille, 1831: 56.—H. Milne Edwards, 1837a: 112.—H. Milne Edwards, 1840: 112. – Bolivar, 1875: 20–21.

Cancer (Pagurus) Carabus : Gmelin and Linne´, 1790: 2984.

Cancer (Astacus) Carabus : Herbst, 1796: 67–68.

Albunea symnista [sic]: Rafinesque­Schmaltz, 1814: 20. – Lucas, 1849a: 27–28 *. – Lucas, 1849b: pl. 3, fig. 2*. – Heller, 1863: 153 (part). – Barrois, 1888: 18–19, 75, 82, 89, 93–94. – Bolivar, 1892: 128 (list). – Ferrer Aledo, 1914: 68. – Miranda y Rivera, 1933a: 22. – Miranda y Rivera, 1933b: 1 (list). – Holthuis, 1954c: 34. – Holthuis, 1956: 228. – ICZN, 1958: 222 (not Albunea symmysta ( Linnaeus, 1758)) .

cancer carabus: Lamarck, 1818: 224 .

Albunea Guerinii Lucas, 1853: 45–47 , pl. 1, fig. 9. – Bolivar, 1875: 20–21. – Carus, 1885: 496. – Rodriguez Femenias, 1887: 5. – de Buen, 1887: 425. – Bolivar, 1916: 251 (list). – de Buen, 1916: 359.

Albunaea [sic] Guerini: Stimpson, 1858: 230 (list).

Albunea sp. Larrinúa y Azcona, 1874: 46.

‘‘ Albunea ?’’ Barceló y Combis, 1875: 63.

Albunea guerinii: Miers, 1878: 327–328 . – Aharoni, 1937: 1136. – Aharoni, 1944: 41 *.

Albunea carabus: Ortmann, 1896: 224 (list). – Ortmann, 1901: 1276. – Monod, 1933: 473 (part). – Gordon, 1938: 186, fig. 3b. – Bouvier, 1940: 179–181, figs. 132, 133. – Holthuis, 1954c: 34. – Piguet, 1955: 14. – Holthuis, 1956: 228. – Monod, 1956: 40–42, figs 10–14. – Holthuis and Gottleib, 1958: 77, 116*. – ICZN, 1958: 222. – Figueira, 1960: 6. – Gauld, 1960: 66 (list). – Zariquiey Alvarez, 1961: 103–109, figs. 1–3. – Crosnier, 1967: 341 *. – Pérés, 1967: 456. – Zariquiey Alvarez, 1968: 294–296, fig. 104. – Moncharmont, 1969: 434–439, figs. 1– 3. – Rubió and Holthuis, 1976: 41–46 *. – Kaestner, 1980: 336, fig. 13–26b. – Riedl, 1983: 488, pl. 179. – Beschin and De Angeli, 1984: 92, fig. 2. – Pretus, 1985–1989a: 258. – Pretus, 1985–1989b: fig. 92. – Coêlho and Calado, 1987: 42–43, table 1. – García Socias and Massuti Jaume, 1987: 79, 91 (list). – García Socias and Gracia, 1988: 55. – Seridji, 1988: 1293– 1298, figs. 1–3. – Guillén Nieto, 1990: 165– 167, fig. 64. – Brunet and Vicente, 1992: 168– 169, figs. 1, 2. – García Raso et al., 1992: 131, 133, 258 (list). – Wirtz and Martins, 1993: 57– 58. – Calado, 1995: 27–30, pl. 4, fig. a, pl. 5, fig. a, pl. 6, figs. a–f*. – González Pérez, 1995: 165, fig. 102. – Giacobbe and Spano`, 1996: 719–725, pl. 1. – Calado, 1997a: 17, 21–22.— Spanò et al., 1999: 617–620. – d’Udekem d’Acoz, 1999: 171. – Boyko, 1999: 161.

Albunea barbara (Lucas ms) Ortmann, 1896: 224 (list) (nomen nudum).

Albunea Guerini : A. Milne Edwards and Bouvier, 1900: 275. – Ferrer y Galdiano, 1918: 413 (list).

Albunea symmista [sic]: Holthuis, 1954c: 34 (not Albunea symmysta ( Linnaeus, 1758)) .

? Albunea sp. B Lebour, 1959: 129 , fig. 15.

Albunea Carabus : Seridji, 1988: 1293, 1298.

Albunea aff. carabus: Fransen, 1991: 48 , 77.

not Hippa caerulea Risso, 1816: 50–51 . – Risso, 1827: 36–37 (= Gnathia sp. cf. phallonajopsis Monod, 1925 ).

not Hippa coerulea [sic]: Risso, 1844: 94. – Hope, 1851: 12 (= Gnathia sp. cf. phallonajopsis Monod, 1925 ).

not Albunea guerinii: Stebbing, 1914: 281 (= Albunea gibbesii Stimpson, 1859 ).

not Albunea carabus: Balss, 1916a: 37 *. – Monod, 1933: 473 (part) (= Albunea elegans A. Milne Edwards and Bouvier, 1898 ).

not Albunea guerinii: Stebbing, 1917: 26 (= Albunea sp. indet.).

not Albunea carabus: Chace, 1966: 635 (= Albunea gibbesii Stimpson, 1859 ).

not Albunea carabus: Holthuis, 1977: 61–62 (= Gnathia sp. cf. phallonajopsis Monod, 1925 ).

MATERIAL EXAMINED: Lebanon: Vicinity of Beirut, coll. M. E. Ericksen: 2 ♀, 16.3– 16.6 mm cl ( USNM 260973).

Israel: Nahr­Rubin, coll. unknown: 1 ♀, 14.7 mm cl ( BMNH 1920.12.16.3) ; Haifa Bay , June 18, 1987, coll. unknown: 1 ♀, 16.5 mm cl ( ZMTAU NS 24665 ) .

Tunisia: Public beach, Raouad, Jan. 30, 1973, coll. J. Moncef and R. B. Manning: 1 Ƌ, 14.0 mm cl ( USNM 256595).

Algeria: Between Lebon and Hussein , 36°44̍N, 03°05̍E, 2–7 m, June–July 1941, coll. H. Nouvel: 1 Ƌ, 4.4 mm cl ( RMNH 34726 ) ; Oran, July 1881, coll. M. Deshayes on ‘‘ Travailleur’ ’: 2 Ƌ, 12.1–13.3 mm cl, 1 ♀, 18.6 mm cl (MNHN­Hi 10) ; Gulf of Oran, June 6, 1900, coll. P. Pallary: 1 ♀, 20.7 mm cl (MNHN­Hi 148) ; Cape Matifou , 1840–1842, M. Deshayes: 1 ♀, 15.5 mm cl, syntype of A. guerinii (MNHN­Hi 189) ; Cape Matifou , 1840–1842, coll. M. Deshayes: 1 Ƌ, 12.4 mm cl, syntype of A. guerinii (MNHN­Hi 190) ; Cape Matifou , 1840–1842, coll. M. Deshayes: 2 Ƌ, 13.5–16.0 mm cl, syntypes of A. guerinii (MNHN­Hi 9) ; Rade de Bône , 15–20 m, 1908, coll. E. Chevraux: 3 ♀, 13.9–18.1 mm cl (MNHN­Hi 1) .

Spain: Off San Andres de Llavaneras , near Arenys de Mar, Barcelona, 4–5 m, Aug. 8, 1973, coll. Tau­Tau: 1 Ƌ, 18.0 mm cl ( RMNH 36273 ), 1 Ƌ, 19.5 mm cl ( AMNH 18077 ex RMNH 36273 ) ; between San Pol de Mar and San Andres de Llavaneres , near Arenys de Mar, Barcelona, 3–6 m, Nov. 22, 1972, March 1973, Aug. 3, 1973, coll. Tau­ Tau: 1 Ƌ, 20.0 mm cl ( RMNH 36274 ), 3 Ƌ, 19.1–21.0 mm cl ( USNM 258373 ex RMNH 36274 ) ; between San Pol de Mar and San Andres de Llavaneres , near Arenys de Mar, Barcelona, 3–6 m, Aug. 27, 1974, coll. Tau­ Tau: 1 Ƌ, 20.3 mm cl ( USNM 260867 ) ; Arenys de Mar , Barcelona, 3–7 m, June 11, 1971, coll. M. Rubio: 1 Ƌ, 18.6 mm cl ( RMNH 28713 ) ; Canet de Mar , Barcelona, 5–7 m, June 23, 1972, coll. Tau­Tau: 1 Ƌ, 18.4 mm cl, 1 ♀, 23.5 mm cl ( RMNH 28911 ) ; Canet de Mar , Barcelona, 5 m, Nov. 1972, coll. M. Rubio: 2 Ƌ, 18.1–19.2 mm cl (MNHN­Hi 85) .

Portugal: Funchal Bay , Madeira, 10–15 m, March 31, 1958, coll. A. Figuera: 1 ♀, 17.7 mm cl ( ZMUC 2712 ) ; Sta. 25, 36°59̍N, 25°03̍W, 10–18 m, May 28, 1981, coll. CANCAP V: 1 oviger, 10.0 mm cl, 1 juvenile, 4.0 mm cl ( RMNH 36282 ) .

Cape Verde: Sta. 7.068, west­southwest of Ilhéu Calheta do Visto, west of Boa Vista, 16°11̍N, 22°59̍W, 40 m, Aug. 27, 1986, coll. CANCAP: 1 ♀, 5.1 mm cl ( RMNH 38605).

Ivory Coast: Sta. 17, Gulf of Guinée, 27 m, June 21, 1956, coll. ‘‘Calypso’’: 8 Ƌ, 9.4– 10.4 mm cl, 6 ♀, 8.3–10.5 mm cl ( MNHN­ Hi 111).

Ghana: Trawl 1, Agassil, coll. R. Bassindale: 1 ♀, 11.4 mm cl ( BMNH 1957.12.4.1).

Togo: Côtes du Togo, 06°07̍N, 01°34̍E, 26 m, Oct. 5, 1963, coll. A. Crosnier: 1 Ƌ, 11.8 mm cl (MNHN­Hi 5). Dahomey: 06°16̍N, 02°26̍E, 22 M, Oct. 9, 1963, coll. A. Crosnier: 1 unsexable, unmeasurable specimen (MNHN­Hi 2); 06°14̍N, 02°26̍E, 30 m, Oct. 9, 1963, coll. A. Crosnier: 1 ♀, 4.9 mm cl (MNHN­Hi 3); 06°09̍20̎N, 01°52̍E, 35 m, Oct. 4, 1963, coll. A. Crosnier: 1 Ƌ, 11.2 mm cl (MNHN­Hi 4); 06°14̍30̎N, 01°48̍E, 15 m, Oct. 4, 1963, coll. A. Crosnier: 1 juvenile, 3.3 mm cl (MNHN­Hi 6).

No Data: 4 Ƌ, 13.0– 16.2 mm cl ( MNHN­ Hi 90); 3 Ƌ, 14.8–15.9 mm cl (MNHN­Hi 91).

DIAGNOSIS: Carapace wider than long, covered with lightly setose crenulate grooves. Anterior margin with 9–14 spines on either side of ocular sinus. Setal field with narrow lateral elements and straight anterior margin. CG1 with separate posterior lateral elements; CG4 with two short and two long medial elements between longer supralateral elements of CG4; CG5 present as two curved, oblique elements; CG6 and CG7 separate; CG8 broken; CG11 present. Rostrum present, not reaching posterior margin of ocular plate. Ocular plate triangular. Distal peduncular segments dorsoventrally flattened and triangular in shape, tapering at tip, approximated along mesial margins at base, lateral margins broadly convex, mesial margins straight. Cornea at tip. Dactylus of pereopod II with heel produced, subquadrate and rounded. Dactylus of pereopod III with heel produced, subquadrate and rounded. Dactylus of pereopod IV sinuous from base to tip, with low rounded heel and shallow indent. Telson of male broadly triangular, as long as wide, with broadly rounded tip; thickly calcified in proximal third and medial strip, inflated dorsally; distolateral two­thirds decalcified, median and transverse rows of thin setae. Telson of female flattened, ovate, and evenly calcified with slightly truncate tip.

DESCRIPTION: Carapace (fig. 87A) wider than long. Anterior margin slightly concave on either side of ocular sinus, becoming convex laterally, with 9–14 large spines (n = 6) along length. Rostrum as small acute tooth, not reaching proximal margin of ocular plate. Ocular sinus smoothly concave, with few small spinules. Frontal region smooth; setal field narrow anteriorly and posteriorly; posterior lateral elements reduced to narrow bands of setae. CG1 parallel to anterior mar­ gin of carapace, sinuous, strongly crenulate, divided into medial fragment and curved, posteriorly displaced lateral elements. Mesogastric region smooth; CG2 absent; CG3 broken into five to eight short elements between posterior lateral elements of CG1; CG4 with two short and two long medial elements between longer supralateral elements of CG4. Hepatic region smooth, with oblique setose groove at median of lateral margin. Epibranchial region generally triangular, smooth; posterolateral margin without rows of setae. Metagastric region smooth; CG5 present as two curved, oblique elements. CG6 strongly crenulate, strongly anteriorly concave medially, and sloping out to anteriorly convex lateral thirds. CG7 oblique, not reaching lateral margins of median segment of CG6. Cardiac region smooth; CG8 present as two short medial elements. CG9 present as two short elements with wide gap at midline. CG10 present as two long elements. CG11 present as two short elements. Post­ CG11 element absent. Branchial region with numerous short, transverse rows of setae. Posterior margin deeply and evenly convex, with submarginal groove reaching halfway up margin of posterior concavity. Branchiostegite with short anterior submarginal spine; anterior region with scattered short, transverse lines ventral to linea anomurica; with many short rows of setae and sparsely covered with long plumose setae ventrally; posterior region membranous, with numerous irregular fragments and sparsely covered with long plumose setae.

Ocular plate (fig. 87B) triangular with deep median indentation; median peduncular segments present as small ovate calcified areas lateral to ocular plate. Distal peduncular segments elongate, subtriangular, with convex lateral and straight mesial margins, cornea covering lateral three­fourths of distal tip; lateral margins with notch just proximal to cornea; mesial margins approximated along length; mesial and lateral margins with long plumose setae; tuft of plumose setae at proximolateral ventral angles and ventromedial row of plumose setae extending from tuft to base of cornea.

Antennule (fig. 87C) with segment III narrow proximally, expanding distally to twice proximal width; with plumose setae on dor­ sal and ventral margins and sparsely scattered on lateral surface; dorsal exopodal flagellum with 86–108 articles (n = 6), long plumose setae on dorsal and ventral margins; ventral endopodal flagellum with three to five articles (n = 6), plumose setae on dorsal and ventral margins. Segment II medially inflated in dorsal view, with plumose setae on dorsal and ventral margins and scattered on ventrolateral third of surface. Segment I wid­ er than long, unarmed; dorsal third of lateral surface rugose with long plumose setae; long plumose setae on dorsal and ventral margins.

Antenna (fig. 87D) with segment V approximately two times longer than wide, with long plumose setae on dorsal and ventral margins and scattered on lateral surface; flagellum with seven articles (n = 6), long plumose setae on dorsal, ventral, and distal margins. Segment IV expanded distally, with long plumose setae on dorsal, ventral, and distal margins, and row of setae on dorsolateral surface. Segment III with long plumose setae on ventral margin and in short row on surface. Segment II short, widening distally, rugose, with plumose setae on margins and scattered on lateral surface; antennal acicle long, thin, and reaching to distal margin of segment IV, with long plumose setae on dorsal margin. Segment I rounded proximally, flattened ventrolaterally, with long plumose setae on margins and scattered on surface rugae behind spine; lateral surface with acute spine dorsodistally (occasionally two spines), with low semicircular dorsolateral lobe ventrodistal to spine; segment with ventromesial antennal gland pore.

Mandible (fig. 87E) incisor process with two teeth; cutting edge with one tooth. Palp three­segmented, with plumose setae on margins and long, thick, simple setae arising from bend in second segment and on distal margin of terminal segment.

Maxillule (fig. 87F) distal endite proximally narrow, widening to inflated distal end, with thick simple setae on distal margin and thin simple setae on dorsal margin. Proximal endite with thick simple setae on distal margin. Endopodal external lobe truncate distally and curled under; internal lobe reduced with three thick setae at distolateral margin.

Maxilla (fig. 87G) exopod evenly rounded, with plumose setae along distal margin. Scaphognathite bluntly angled on posterior lobe, with plumose setae.

Maxilliped I (fig. 87H) epipod with plumose setae on margins, distolateral surface, and mesial surface. Endite tapered distally and subequal to first segment of exopod. Exopod with two segments: proximal segment narrow, margins parallel with plumose setae; distal segment spatulate, longer than wide, broadest medially, margins and mesioventral surface with long plumose setae. Endopod flattened and elongate, reaching two­thirds of distance to distal end of proximal exopodal segment; plumose setae on margins and median of lateral surface.

Maxilliped II (fig. 87I) dactylus evenly rounded, length equal to width, with thick simple setae distally and on distolateral surface. Propodus two times wider than long, slightly produced at dorsodistal angle, with plumose setae on dorsal margin and patch of long simple setae on lateral surface and ventrolateral angle. Carpus not produced dorsodistally, approximately two times longer than wide; long simple setae on dorsal margin and in short row on lateral surface. Merus approximately three times longer than wide, margins parallel; with simple and plumose setae on margins and scattered on surface. Basis­ischium incompletely fused with plumose setae on margins. Exopod one­third longer than merus, flagellum with one elongate article, longer than carpus.

Maxilliped III (fig. 87J) dactylus with rounded tip; long plumose setae on margins and lateral surface. Propodus dorsodistally inflated, with longitudinal median row of plumose setae on lateral surface; margins with plumose setae. Carpus produced onto propodus almost one­half length of propodus; lateral surface with two rows of plumose setae; plumose setae on margins. Merus inflated, unarmed, with plumose setae on margins and scattered on lateral surface. Basisischium incompletely fused, without crista dentata. Exopod two­segmented: proximal segment small; distal segment styliform, tapering, approximately one­third length of merus; with plumose setae on margins; without flagellum.

Pereopod I (fig. 88A) dactylus curved and tapering; lateral and mesial surfaces smooth; dorsal margin with few rugae proximally and long plumose and short simple setae; ventral margin with short simple setae. Propodal lateral surface with numerous short, transverse rows of setose rugae; dorsal margin unarmed; ventral margin distally produced into acute spine; cutting edge lacking teeth, lined with long plumose setae; dorsal margin with long plumose setae, ventral margin with short simple setae. Carpus with dorsodistal angle produced into strong corneous­tipped spine; dorsal margin with short transverse grooves behind spine; dorsal and distal margins with short plumose setae; lateral surface with small distal rugose area, with few transverse setose ridges on distal half of surface; mesial surface smooth, with medial trans­ verse and subdorsal row of setae, margins with long plumose setae. Merus unarmed; lateral surface with scattered transverse rows of long plumose setae, margins with long plumose setae; mesial side with few scattered setae; fully calcified. Basis­ischium incompletely fused, unarmed. Coxa unarmed.

Pereopod II (fig. 88B) dactylus smooth; base to heel slightly concave, heel rounded and not produced, heel to tip with narrow, acute indent, tip acute, tip to base broadly convex; lateral surface smooth, with several small tufts of short setae in roughly straight line across medioproximal surface, several widely spaced submarginal tufts of short setae dorsodistally; mesial surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae, with patch of long plumose setae at base. Propodal dorsal surface smooth, ventral margin inflated and rounded; oblique row of long plumose setae on distal margin of lateral surface; distal and ventral margins with long plumose setae; dorsolateral surface as narrow, oblique, flattened shelf, with short setae on dorsal margin and long plumose setae on ventral margin; mesial surface with elevated, curved setose ridge from ventral junction with dactylus almost to ventral proximal junction with carpus. Carpus strongly produced and rounded dorsodistally, dorsal margin smooth proximally, serrate distally; lateral surface smooth, with small setose mat at tip of produced area and irregular, broken row of rugae and submarginal elevated ridge ventrally, rugae and ridge with long plumose setae; margins with long plumose setae; mesial surface smooth with row of long plumose setae distally, ventrally, and subdorsally. Merus with large median decalcified window covering nearly all of lateral surface, with few scattered long plumose setae on surface and margins; mesial surface nearly smooth with two long rows of setae. Basisischium incompletely fused and unarmed. Coxa unarmed.

Pereopod III (fig. 88C) dactylus with base to heel slightly concave, heel rounded and not produced, heel to tip with broadly concave indent, tip acute, tip to base smoothly convex; lateral surface smooth, with several small tufts of short setae in roughly straight line across medioproximal surface, dorsodistal margin with tufts of short setae; ventral margin with long plumose setae, dorsal margin with short simple and plumose setae; mesial surface smooth with plumose setae proximally at junction with propodus. Propodus not inflated dorsoventrally; lateral surface smooth, with long plumose setae in oblique row, simple setae on dorsal margin; dorsolateral surface narrow, oblique, flattened; mesial surface smooth. Carpus produced dorsodistally, exceeding proximal margin of propodus by one­half length of propodus; dorsolateral margin unarmed; lateral surface slightly rugose dorsodistally, with mat of short setae and two interrupted rows of setae ventrally; mesial surface smooth, with long plumose setae on distal margin and in row on distal half of surface. Merus smooth, with large decalcified window covering nearly half of lateral surface medially; dorsal and ventral margins unarmed, with long plumose setae; distolateral margin with long plumose setae; mesial surface smooth. Basis­ischium incompletely fused and unarmed. Coxa unarmed. Female with large gonopore on anterior mesial margin of coxa, surrounded with short plumose setae; male without pore.

Pereopod IV (fig. 88D) dactylus with base to tip convex proximally, concave distally, with shallow indent, tip acute, tip to base convex; lateral surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae; mesial surface with dorsal decalcified region, demarcated ventrally by longitudinal elevated ridge with row of short setae; with setose punctations ventral to decalcified window. Propodus expanded dorsally and ventrally; ventral expansion reaching ventral margin of dactylus, margin with long plumose setae; dorsal expansion with row of long plumose setae dorsally, oblique area smooth; lateral and mesial surfaces smooth. Carpus slightly produced dorsodistally; ventral three­fourths of lateral surface and mesial surface smooth, with few scattered setae, dorsodistal quarter of lateral surface with mat of short setae; dorsal margin with short simple and long plumose setae; ventral margin with short simple setae; mesial surface decalcified medially. Merus lateral surface with scattered short transverse rows of setae, dorsal and ventrodistal margins with long plumose setae; proximoventral half of mesial surface with large decalcified window. Basis­ischium incompletely fused and unarmed. Coxa unarmed.

Abdomen (fig. 88E) with somite I length and width subequal, widest posteriorly; dorsal surface with anterior margin straight; posterior margin curved, with elevated submarginal row of short setae; small transverse decalcified windows laterad of segment medi­ an. Somite II dorsal surface with submarginal transverse ridge anteriorly; with small transverse decalcified windows laterad of segment median just anterior to submarginal ridge; pleura expanded and directed anterolaterally; lateral margins angled, anterior and lateral margins with long plumose setae, posterior margin with short setae; posteromesial angle with mat of short simple setae. Somite III similar to somite II, but narrower, shorter; pleura thinner and shorter than on somite II, directed posterolaterally but with distal tips swept forward, with setae as in somite II; anterolateral angle subacute; dorsal surface obliquely flattened anterolaterally. Somite IV similar to somite III, but thinner and shorter; dorsal surface with few short setae anterolaterally; pleura thinner and shorter than on somite III, directed posterolaterally with distal tips slightly swept forward; dorsal surface obliquely flattened anterolaterally; margins with long plumose setae. Somite V wider than somite IV; lateral margins with plumose setae; pleura absent. Somite VI slightly broader than somite V; dorsal surface and distal margin with short transverse rows of setae laterad of midline; pleura absent.

Females with uniramous, paired pleopods on somites II–V; males without pleopods. Telson of male (fig. 88F) broadly triangular, as long as wide, with broadly rounded tip; thickly calcified in proximal third and medial strip, inflated dorsally; distolateral two­thirds decalcified; median longitudinal groove extending one­half length, rows of long simple setae demarcating juncture of calcified and decalcified regions and in tuft at distal end; proximolateral angles without setae; margins with long simple setae. Telson of female (fig. 88G) flattened, ovate, and evenly calcified with slightly truncate tip; median groove similar to male, row of short simple setae on either side of median groove from midpoint of telson continuing distally to three­fourths length of telson and in short transverse submarginal lateral rows approximately onefourth distal from proximal margin; proximolateral angle without setae, margins with long simple setae.

DISTRIBUTION: From Lebanon westward through the Mediterranean and from the Azores and Madeira southward to Dahomey, in up to 40 m depth.

MAXIMUM SIZE: Males: 21.0 mm cl; females: 23.5 mm cl.

TYPE SPECIMENS: The type(s) of Cancer carabus are lost. MNHN­Hi 189 (syntype of A. guerinii ), MNHN­Hi 190 (syntype of A. guerinii ), MNHN­Hi 9 (2 syntypes of A. guerinii ). Lucas (1849a, 1853) cited 16 total specimens (= syntypes) of A. guerinii . Only four specimens in the MNHN are labeled as syntypes and possess the correct data. It may be that MNHN­Hi 90 and MNHN­Hi 91 are also syntypes of A. guerinii , but they have no data associated with them that would confirm this.

TYPE LOCALITIES: Cancer carabus : ‘‘M. Mediterraneo’’ ( Linnaeus, 1758). As pointed out by Rubió and Holthuis (1976), the collector of the type (s), Erik Brander, was stationed in Algeria, and this is probably where the type (s) came from, although it cannot be considered the type locality in the absence of a primary type specimen. Albunea guerinii : Algeria.

REMARKS: At least part of the difficulty early authors had in identifying this taxon as an Albunea was caused by Linnaeus’ (1758) placement of the species within Cancer . Cancer Carabus (Linnaean no. 48) was separated by eight intervening taxa from Cancer Symmysta (Linnaean no. 39), the only other albuneid described by Linnaeus (1758). These eight intermediate taxa were an assortment of anomurans, crayfish, and shrimp which, by themselves, offered no clue as to the relationships of carabus and symmysta . By 1767, Linnaeus had moved the two taxa closer together, to no. 68 and no. 70 on his list, and separated them by a galatheid. The ‘‘rostrum dentibus 2 parallelis mobilibus depressis’’ described by Linnaeus (1758) for this species is actually the ocular peduncles rather than the rostrum. This was probably the key morphological feature that caused confusion as to what sort of crustacean Linnaeus was referring to ( Miers, 1878). Linnaeus (1758) cited this species as ‘‘Testa magnitudine extimi pollicis’’ (as long as the last joint of the thumb), and this has been extrapolated to a carapace length of approximately 17 mm (Rubió and Holthuis, 1976), although larger specimens are known. Calado (1995) erroneously cited BMNH 1920.12.16.3 as the ‘‘holótipo.’’ No Linnaean material of this species exists, no neotype has ever been designated, and no neotype is needed, as this species is the only albuneid in the Mediterranean and is only distantly related to the other albuneid species in the east Atlantic. The species has been called ‘‘Leeuw­Krab’’ ( Houttuyn, 1769), ‘‘Kahn Krabbe’’ ( Statius Müller, 1775), and ‘‘Löw­ enkrebs’’ (Herbst, 1796), but no general common name has been adopted.

Lucas (1853), in describing A. guerinii , was the first to recognize and discuss the sexual dimorphism in telson shape within the genus. Zariquiey Alvarez’s (1961: fig. 1) illustration of this species is one of the most accurate drawings published for any albuneid before 1999 (fig. 1C, herein), and his illustrations of the male and female telsons are another rare example of recognition of the sexual dimorphism present in this genus.

Brunet and Vicente (1992) incorrectly cit­ ed Monod as the first author to unequivocally assign this taxon to Albunea ; that assignation was first indicated by Ortmann (1896). Ortmann (1896) also introduced the manuscript name ‘‘ Albunea barbara Lucas’’ in synonymy with A. carabus . That name is both a nomen nudum and an unavailable name. The specimen cited from ANSP by Ortmann (1896) and identified as A. barbara Lucas is not extant (Boyko, personal obs.).

Holthuis’s (1977) placement of Hippa caerulea Risso in synonymy with A. carabus is incorrect, as Hippa caerulea is actually a gnathiid isopod (see appendix 2).

Although there are repeated statements in the literature that this species is ‘‘muy raro’’ ( Zariquiey Alvarez, 1968; Brunet and Vicente, 1992), it is likely more common in its preferred sand bottom habitats than current data suggest. The apparent rarity of this species in the eastern basin of the Mediterranean is likely due to limited collecting efforts rath­ er than to unspecified ‘‘ecological factors’’ (see Spanò et al., 1999). The species is also probably more widely distributed in the eastern basin than current data indicate. In fact, A. carabus is occasionally washed ashore in large numbers, possibly due to mortality from freshwater runoff ( Piguet, 1955). This is apparently the only species of albuneid in the Azores, but is only known from a few specimens from that location ( Figueira, 1960).

Seridji (1988) described zoeal stages I, II, and III from plankton collected off Algeria. Seridji (1988) speculated that A. carabus might have the same number of zoeal stages (five) as A. symmysta , but as these two species are not particularly closely related and larval development in the genus is overall poorly known, this seems a premature conclusion. The west African larvae reported by Lebour (1959) as ‘‘ Albunea sp. B’’ resemble the early (stage II or III) zoeae of A. carabus in the shape of the posterior telson margin. The coloration of adults is: carapace dark purple with brownish areas, dorsal regions of pereopods and abdomen purple, antennular flagellae ringed with purple and yellowish brown (Rubió and Holthuis, 1976).

This species typifies the ‘‘ carabus ­group’’ of Albunea , which also includes A. danai and A. bulla . It is the sister species to A. danai .

Albunea asymmetrica (P. Müller, 1979) Figure 89

Mioranina asymmetrica P. Müller, 1979: 278–279 , pls. 9, 10*.

Albunea asymmetrica: P. Müller, 1984: 62 , pl. 28, figs. 4–6*.

MATERIAL EXAMINED: Hungary: MGy–2– 1, Upper Badenian , Budapest, Gyakorlóút, coll. P. Müller: 1 carapace fragment, 10.5 × 13.6 mm (l × w), holotype (plaster cast of HNHM) ; MGy–2–2, Upper Badenian, Budapest, Gyakorlóút , coll. P. Müller: 1 carapace fragment, 13.8 × 12.2 mm (l × w), paratype (plaster cast of HNHM; pl. 9, fig. 3 in P. Müller, 1979) ; MGy–2–2, Upper Badenian, Budapest, Gyakorlóút , coll. P. Müller: 1? merus, 13.0 × 8.0 mm (l × w), paratype (plaster cast of HNHM; pl. 9, fig. 2 in P. Müller, 1979) ; MGy–2–2, Upper Badenian, Budapest, Gyakorlóút , coll. P. Müller: 1 carapace fragment, 12.5 × 9.5 mm (l × w), paratype (plaster cast of HNHM; pl. 10, fig. 2 in P. Müller, 1979) ; MGy–2, Upper Badenian, Budapest, Gyakorlóút , coll. P. Müller: 1 carapace fragment, 20.2 × 10.1 mm (l × w), paratype (latex cast of HNHM; pl. 10, fig. 1 in P. Müller, 1979) ; MGy–2, Upper Badenian , Budapest, Gyakorlóút, coll. P. Müller: 6 fragments ( HNHM) ; MR8–16 , Upper Badenian, Budapest­Rákos, Layer 8, coll. P. Müller: 2 fragments, 9.5 × 9.8 mm, 7.5 × 8.0 mm (l × w) ( HNHM) .

DIAGNOSIS: Carapace as long as wide, covered with crenulate grooves. Anterior margin with 10 spines on either side of ocular sinus. Setal field with thin lateral elements and straight anterior margin. CG1 with separate posterior lateral elements; CG4 with two long medial elements between longer supralateral elements of CG4; CG5 unknown; CG6 and CG7 fused; CG8 unknown; CG11 unknown. Rostrum present.

DESCRIPTION: Carapace (fig. 89A–C) approximately as wide as long. Anterior margin slightly concave on either side of ocular sinus, becoming convex laterally with 10 large spines (n = 1) along length. Rostrum as small acute tooth. Ocular sinus smoothly concave and unarmed. Frontal region smooth; setal field narrow anteriorly and widening posteriorly; posterior lateral elements reduced to narrow bands of setae. CG1 parallel to anterior margin of carapace, sinuous, strongly crenulate, divided into medial fragment and curved, posteriorly displaced lateral elements. Mesogastric region smooth; CG2 presence unknown; CG3 present but composition unknown; CG4 with two long medial elements between longer supralateral elements of CG4. Hepatic region smooth, with oblique setose groove at median of lateral margin. Epibranchial region generally triangular, smooth. Metagastric region smooth; CG5 presence unknown. CG6 strongly crenulate, strongly anteriorly concave medially and sloping out to anteriorly convex lateral thirds. CG7 oblique, not reaching lateral margins of median segment of CG6. Cardiac region unknown; CG8 unknown. CG9 unknown. CG10 unknown. CG11 unknown. Branchial region with numerous short, transverse rows of setae. Branchiostegite (fig. 89D) with short anterior submarginal spine; anterior region with scattered rugae.

Pereopod I subchelate. Dactylus unknown. Propodal lateral surface with numerous short, transverse rows of setose rugae; dorsal margin unarmed.

DISTRIBUTION: Known only from the Upper Badenian formation, Budapest, Hungary.

TYPE SPECIMENS: HNHM MGy–2–1 (holotype), HNHM MGy–2–2 (8 paratypes).

TYPE LOCALITY: MGy–2–1, Upper Badenian, Budapest, Gyakorlóút, Hungary.

REMARKS: The species was first described as the type of the ‘‘raninid’’ genus Mioranina P. Müller, 1979 , illustrating once again the convergent morphology between raninds and albuneids. None of the material of this taxon is well preserved, especially when compared to other known albuneid fossils (e.g., Beschin and De Angeli, 1984). This material is quite similar to A. carabus in its carapace grooves, shortness of branchiostegite spine, and proportions of length and width. The number and size of the anterior carapace margin spines of A. asymmetrica are not appreciably different from A. carabus , contrary to the statements of P. Müller (1984). The relatively recent age (15 mybp, Tertiary: Miocene) of this fossil material also suggests that it may be conspecific with an extant taxon. Additional better preserved material is needed to determine if A. asymmetrica and A. carabus are synonymous or closely related species.

Albunea danai Boyko, 1999 Figures 90 , 91

Albunea danai Boyko, 1999: 145 (list), 155–161, figs. 5, 6*.

MATERIAL EXAMINED: USA: Hawaii: Halonu Blow Hole dive site, south shore, Oahu , 12.2–13.7 m, April 4, 1997, coll. R. Holcom: 1 Ƌ, 16.7 mm cl, holotype ( QM W23105) ; Kailua, Oahu , March 1938, coll. unknown: 1 ♀, 16.8 mm cl, allotype ( BPBM S11782 ) ; off Eaa Beach, near Pearl Harbour , 15 fms (= 27.4 m), July 5, 1964, coll. B. R. Wilson on ‘‘ Pele’ ’: 1 ♀, 10.8 mm cl, paratype ( WAM 10422) ; off Waikiki, Oahu , 20 ft (= 6.1 m), May 30, 1948, coll. Allen and V. Smith: 1 ♀, 10.8 mm cl, paratype ( AMNH 17716 ex BPBM S5348 ) ; off Sand Island, Oahu , 16– 25 fms (= 29.3–45.7 m), July 17, 1959, coll. Pele Expedition: 1 Ƌ, 10.4 mm cl, paratype ( AMNH 17717 ex BPBM S6776 ) ; off Honolulu , 15–22 fms (= 27.4–40.2 m), Feb.– March 1962, coll. T. Richert: 1 Ƌ, 16.4 mm cl, 1 ♀, 15.5 mm cl, paratypes ( WAM 23390) ; Waikiki, Oahu , 75 ft (= 22.7 m), May 23, 1948, coll. Smith and Allen: 1 Ƌ, 11.6 mm cl, paratype ( BPBM S5343 ) ; Kahana Bay, Oahu , 25–30 fms (= 45.7–54.9 m), July 25, 1959, coll. Pele Expedition: 1 Ƌ, 8.6 mm cl, paratype ( BPBM S6775 ) ; Diamond Head, Oahu , 25–45 fms (= 45.7–82.3 m), Sept. 9, 1959, coll. Pele Expedition: 1 ♀, 13.0 mm cl, paratype ( BPBM S6777 ) ; Kanoehe Bay, Oahu , 1924, coll. unknown: 1 Ƌ, 4.0 mm cl ( BPBM S7806 ) .

DIAGNOSIS: Carapace slightly longer than wide, covered with lightly setose grooves. Anterior margin with eight or nine spines. Setal field with narrow lateral elements and slightly concave anterior margin; posterior lateral elements not extending to posterior lateral elements of CG1. CG1 with separate posterior lateral elements; CG4 with two to four short medial elements; CG5 divided into two lateral elements, not nearly reaching margins of CG6; CG6 and CG7 separate, but almost approximate; CG8 with one or two posteriorly displaced median elements separated from lateral elements; CG11 absent. Rostrum present, not reaching proximal margin of ocular plate. Ocular plate subquadrate. Distal peduncular segments dorsoventrally flattened and elongate, pointed at tip, approximate along mesial margins; lateral margins convex; mesial margins straight proximally, convex distally. Cornea at lateral margin of tip. Antennule with 87–92 flagellar exopodal and three or four endopodal articles. Antenna with seven flagellar articles; acute spine on dorsolateral surface of peduncle segment I. Dactyli of pereopods II and III with heels low and smoothly rounded. Coxa of pereopod III of males with small pore. Telson of male triangular, dorsoventrally flattened laterally and distally, inflated medially. Telson of female flattened, rounded at tip.

DESCRIPTION: Carapace (fig. 90A) slightly wider than long. Anterior margin slightly concave on either side of ocular sinus, becoming convex laterally with eight or nine large spines along length. Rostrum as small acute tooth, extending only one­half of the distance between distal margin of the ocular sinus and ocular plate. Ocular sinus smoothly concave and unarmed. Frontal region smooth; setal field broad posteriorly, narrowing anteriorly, with narrow anterior lateral elements and slightly concave anterior margin; posterior lateral elements thin and not reaching to posterior lateral elements of CG1. CG1 parallel to anterior margin of carapace, faintly sinuous, strongly crenulate, divided into medial fragment and curved, posteriorly displaced lateral elements. Mesogastric re­ gion smooth; CG2 absent; CG3 broken into six short elements approximately equally spaced between posterior lateral elements of CG1; CG4 with two to four short medial el­ ements spaced approximately equally between longer lateral elements of CG4. Hepatic region smooth, with oblique setose groove at median of lateral margin. Epibran­ chial region generally triangular, smooth; posterolateral margin with two short rows of setae. Metagastric region smooth; CG5 divided into two short lateral elements. CG6 strongly crenulate, strongly anteriorly concave medially and sloping out to anteriorly convex lateral thirds. Cervical groove strongly concave at median and sloping out to irregularly convex lateral thirds; median and lateral thirds separated by short setae­free gap lateral to small depressions. CG7 oblique, almost reaching lateral margins of median segment of CG6. Cardiac region smooth; CG8 present as two very short lateromedial elements displaced posteriorly from longer lateral elements. CG9 present as two short lateral grooves with gap at midline. CG10 present as two curved lateral fragments with gap between fragments approximately equal to length of single fragment. CG11 absent. Branchial region with numerous short, transverse rows of setae. Posterior margin deeply and evenly convex, with submarginal groove reaching approximately halfway up either side of posterior concavity. Branchiostegite with short anterior submarginal; anterior region with scattered, short, transverse lines ventral to linea anomurica; with many short rows of setae and sparsely covered with long plumose setae ventrally; posterior region membranous, with numerous irregular fragments and sparsely covered with long plumose setae.

Ocular plate (fig. 90B) subquadrate with median indentation; median peduncular segments reduced to small rounded calcified area on either side of ocular plate. Distal peduncular segments elongate, with distally convex lateral margins, tapering to rounded distal cornea located in lateral notch; mesial margins approximated almost all of length; mesial and proximolateral margins of segment with sparse row of long plumose setae; tuft of plumose setae at proximolateral ventral angle.

Antennule (fig. 90C) with segment III narrow proximally, expanding distally to two times proximal width; with plumose setae on dorsal and ventral margins and sparsely scattered on lateral surface; dorsal exopodal flagellum with 87–92 articles, long plumose setae on dorsal and ventral margins; ventral endopodal flagellum short with three or four articles and plumose setae on dorsal and ventral margins. Segment II medially inflated in dorsal view, with plumose setae on dorsal and ventral margins and scattered on ventrolateral third of surface. Segment I wider than long, unarmed; dorsal third of lateral surface rugose with long plumose setae; long plumose setae on dorsal and ventral margins.

Antenna (fig. 90D) with segment V approximately two times longer than wide, with long plumose setae on dorsal margin and scattered on distal half of lateral surface; flagellum with seven articles, long plumose setae on dorsal, ventral, and distal margins. Segment IV expanded distally, with long plumose setae on dorsal, ventral, and distal margins and two rows of setae on dorsolateral surface. Segment III with long plumose setae on dorsal and ventral margin. Segment II short, widening distally, with plumose setae on margins and scattered on lateral surface; antennal acicle long, thin and exceeding distal margin of segment IV by one­fourth of length of segment IV, with long plumose setae on dorsal margin. Segment I rounded proximally, flattened ventrolaterally with long plumose setae on margins; lateral surface with acute spine dorsally, with low semicircular dorsolateral lobe ventrodistal to spine; segment with ventromesial antennal gland pore.

Mandible (fig. 90E) incisor process with one tooth; cutting edge with one tooth. Palp three­segmented, with plumose setae on margins and long, thick, simple setae arising from bend in second segment.

Maxillule (fig. 90F) distal endite proximally narrow, widening to inflated distal end, with thick simple setae on distal margin. Proximal endite with thick simple setae on distal margin. Endopodal external lobe truncate distally and curled under; internal lobe reduced, with three thick setae at distolateral margin.

Maxilla (fig. 90G) exopod evenly rounded, with plumose setae along distal margin. Scaphognathite bluntly angled on posterior lobe, with plumose setae.

Maxilliped I (fig. 90H) epipod with plumose setae on margins, distolateral surface, and mesial surface (epipod shown curled). Endite tapered distally and subequal to first segment of exopod. Exopod with two seg­ ments; proximal segment narrow, margins parallel, with plumose setae; distal segment spatulate, approximately as long as wide, broadest medially, margins and mesioventral surface with long plumose setae. Endopod flattened and elongate, reaching to distal end of proximal exopodal segment; plumose setae on margins and median of lateral surface.

Maxilliped II (fig. 90I) dactylus evenly rounded, length equal to width, with thick simple setae distally and on distolateral surface. Propodus two times wider than long, slightly produced at dorsodistal angle, with plumose setae on dorsal margin and long simple setae on dorsodistal margin. Carpus not produced dorsodistally, approximately two times longer than wide; long simple setae on dorsal and distal margins. Merus approximately three times longer than wide, margins parallel; with simple setae on ventrolateral margin and plumose setae on dorsolateral margin. Basis­ischium incompletely fused with plumose setae on margins. Exopod one­third longer than merus, with flagellum one­segmented. Maxilliped III (fig. 90J) dactylus with rounded tip; long plumose setae on margins and lateral surface. Propodus with longitudinal median row of plumose setae on lateral surface; margins with plumose setae. Carpus slightly produced onto propodus; lateral surface with row of plumose setae ventromedially; plumose setae on margins. Merus unarmed, with plumose setae on margins and scattered on lateral surface. Basis­ischium incompletely fused, with weak crista dentata of approximately two teeth. Exopod two­segmented: proximal segment small; distal segment styliform, tapering, approximately one­third length of merus; with plumose setae on margins; without flagellum.

Pereopod I (fig. 91A) dactylus curved and tapering; lateral and mesial surfaces smooth; dorsal margin with long plumose and short simple setae; ventral margin with short simple setae. Propodal lateral surface with numerous short, transverse rows of setose rugae; dorsal margin unarmed; ventral margin distally produced into acute spine; cutting edge lacking teeth, lined with long plumose setae; dorsal margin with long plumose setae, ventral margin with short simple setae. Carpus with dorsodistal angle produced into strong corneous­tipped spine; dorsal margin otherwise unarmed; dorsal and distal margins with long plumose setae; lateral surface with small distal rugose area, with few transverse setose ridges on distal half of surface; mesial surface smooth, with few median rows of setae, margins with long plumose setae. Merus unarmed; lateral surface with scattered transverse rows of long plumose setae, margins with long plumose setae; mesial side with few short rows of setae; proximal quarter of mesial surface with decalcified window. Basis­ischium incompletely fused, unarmed. Coxa unarmed.

Pereopod II (fig. 91B) dactylus smooth; base to heel straight, heel smoothly rounded, heel to tip with wide, acute indent, tip acute, tip to base broadly convex distally and concave proximally; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, several widely spaced submarginal tufts of short setae dorsodistally; mesial surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae, patch of long plumose setae at base. Propodal dorsal surface smooth, with ventral margin inflated and rounded; oblique row of long plumose setae on distal margin of lateral surface; distal and ventral margin with long plumose setae; dorsolateral surface as narrow, oblique, flattened shelf, with short setae on dorsal margin and long plumose setae on ventral margin; mesial surface with elevated, curved, setose ridge from ventral junction with dactylus almost to ventral proximal junction with carpus; decalcified region just distal to junction with carpus. Carpus slightly produced and gently rounded dorsodistally, dorsal margin unarmed; lateral surface smooth, with irregular, interrupted row of rugae and submarginal elevated ridge ventrally, rugae and ridge with long plumose setae; margins with long plumose setae; mesial surface smooth, with long plumose setae in scattered patches on dorsal half of surface and on margins. Merus with large median decalcified window covering nearly all of lateral surface, with few scattered setae on surface and margins; mesial surface nearly smooth, with few setae, decalcified area on proximal quarter near junction with basis­ischium. Basis­ischium incompletely fused and unarmed. Coxa with one small spine on anterior margin.

Pereopod III (fig. 91C) dactylus with base to heel straight, heel broadly rounded and low, heel to tip with broadly concave indent, tip acute, tip to base smoothly convex distally to straight proximally; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, dorsodistal margin with tufts of short setae; ventral margin with long plumose setae, dorsal margin with short simple and plumose setae; mesial surface smooth, with plumose setae proximally at junction with propodus. Propodus not inflat­ ed dorsoventrally; lateral surface smooth, with long plumose setae distally, simple setae on dorsal margins; dorsolateral surface narrow, oblique, flattened; mesial surface with scattered long setae on and near distal margin, with decalcified window near junction with carpus. Carpus produced dorsodistally, exceeding proximal margin of propodus by approximately one­third length of propodus, rounded; dorsolateral margin unarmed; lateral surface slightly rugose dorsodistally, with mat of short setae and row of setae ventrally; mesial surface smooth, with long plumose setae on margins and scattered on surface. Merus smooth, with large decalcified window covering nearly half of lateral surface medially; dorsal and ventral margins unarmed, with long plumose setae; laterodistal margin with long plumose setae; mesial surface smooth, with decalcified window at junction with basis­ischium. Basis­ischium incompletely fused and unarmed. Coxa with tubercle on anterior margin. Female with large gonopore on anterior mesial margin of coxa, surrounded with short plumose setae; male with small pore.

Pereopod IV (fig. 91D) dactylus with base to tip convex to concave, tip acute, tip to base straight distally, becoming convex proximally; lateral surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae; mesial surface with dorsal decalcified region, demarcated ventrally by longitudinal elevated ridge with row of short setae; with setose punctations ventral to decalcified window. Propodus expanded dorsally and ventrally; ventral expansion exceeds ventral margin of dactylus, margin with long plumose setae; dorsal expansion with row of long plumose setae medially; lateral and mesial surfaces smooth. Carpus not produced dorsodistally; ventral three­fourths of lateral surface and mesial surface smooth, dorsal fourth of lateral surface with mat of short setae; dorsal margin with short simple and long plumose setae; ventral margin with short simple setae. Merus with scattered, short, transverse rows of setae on lateral surface, dorsal and ventrodistal margins with long plumose setae; mesial surface with large decalcified window proximoventrally. Basis­ ischium incompletely fused and unarmed. Coxa unarmed.

Abdomen (fig. 91E) with somite I approximately as long as wide, widest posteriorly; dorsal surface with anterior margin straight; posterior margin straight with elevated submarginal row of short setae; with small transverse, decalcified windows laterad of segment median. Somite II dorsal surface with submarginal transverse ridge anteriorly; with small transverse decalcified windows laterad of segment median just anterior to submarginal ridge; with tuft of setae at posterolateral angle, extending onto pleura posteromesially; pleura expanded and directed slightly anteriorly; lateral margins rounded, anterior and lateral margins with long plumose setae, posterior margin with short setae. Somite III similar to somite II, but narrower, shorter, and lacking anterior submarginal ridge; small tuft of short thick setae on posterolateral angle; pleura thinner and shorter than on somite II, directed posterolaterally, with setae as in somite II; anterolateral angle acute; dorsal surface obliquely flattened anterolaterally. Somite IV similar to somite III, but thinner and shorter; dorsal surface with few thick setae posterolaterally; pleura thinner and short­ er than on somite III, directed posterolaterally; dorsal surface obliquely flattened anterolaterally; margins with long plumose setae. Somite V wider than somite IV; lateral margins with plumose setae; pleura absent. Somite VI subequal to somite V in length but wider; dorsal surface with short transverse rows of setae laterad of midline anteriorly and posteriorly; lateral margins with long plumose setae; pleura absent.

Females with uniramous, paired pleopods on somites II–V; males without pleopods.

Telson of male (fig. 91F) triangular, slightly longer than wide, with smoothly rounded tip; proximal half heavily calcified, distal half weakly calcified except for large median region; median longitudinal groove extending to distal end of calcified area, line with long, thin, simple setae; junction of proximal and distal regions demarcated by strong line of long setae laterally; calcified plate slightly elevated medially but without ridge. Telson of female (fig. 91G) ovate, longer than wide, rounded distally; dorsal surface smooth, with median longitudinal groove anteriorly; with row of setose punctae lateral to midline from posterior end of longitudinal groove to threefourths of length of telson; margins with long plumose setae.

DISTRIBUTION: Known only from Oahu, Hawaii, USA, 4.8–40.2 m depth.

MAXIMUM SIZE: Males: 16.7 mm cl; females: 16.8 mm cl.

TYPE SPECIMENS: QM W23105 (holotype), BPBM S11782 (allotype), WAM 10422 (paratype), AMNH 17716 (paratype), AMNH 17717 (paratype), WAM 23390 (2 paratypes), BPBM S5343 (paratype), BPBM S6775 (paratype), BPBM S6777 (paratype).

TYPE LOCALITY: Halonu Blow Hole dive site, south shore, Oahu, Hawaii, USA, 12.2– 13.7 m.

REMARKS: The coloration of this species is brownish with reddish­brown setae in life (from color transparencies), and a uniform off­white to tan in preservative.

Based on all available data, this species appears to be a true Hawaiian endemic. However, this conclusion should be accepted cautiously, given the ‘‘endemic’’ label applied to A. speciosa prior to the findings of Boyko (1999). The holotype of A. danai was collected with specimens of A. speciosa (QM W22285) and the two species are sympatric in Hawaii in at least part of their bathymetric ranges.

This species belongs to the ‘‘ carabus ­ group’’ and is the sister species to A. carabus , from the Mediterranean and western Africa, and shares with it a similar shape of the dactyli of pereopods II–IV and telson morphology. Albunea carabus can be easily separated from A. danai by its CG8 of four medial elements, more strongly crenulated CGs, more pronounced heel on the dactyli of pereopods II and III, and a less inflated merus of maxilliped III. Albunea danai can be distinguished from other Indo­West Pacific species, except A. bulla , by the triangular shape of the telson of the male, the rounded dactylus of pereopod III, and setal patterns on the carapace of both sexes. Albunea danai can be separated from A. bulla , which is the sister taxon to the clade containing A. danai and A. carabus , by the different pattern of carapace grooves and the distinctly ‘‘buttonlike’’ morphology of the telson of the males of A. bulla .

Albunea marquisiana Boyko, 2000 Figures 92 , 93

Albunea sp. cf. symnysta [sic] Poupin, 1996b: 26, pl. 12, fig. c*. – Poupin, 1998: 39 *.

Albunea sp. Tudge et al., 1999: 2–5, figs. 1–3*.

Albunea marquisiana Boyko, 2000a: 109–115 , figs. 2, 3*.

MATERIAL EXAMINED: Marquises Islands : Sta. TH X, Haul 1, Haava Straits between Île Tahuata and Île Hiva Oa, 09°52̍S, 139°04̍W, 40 fms (= 73 m), Oct. 1, 1967, coll. D. M. Devaney for National Geographic Society – Smithsonian­Bishop Museum Marquesas Expedition: 1 Ƌ, 10.2 mm cl, holotype ( USNM 268577 ) ; Sta. DW 1279, Île Eiao , 07°59.4̍S, 140°42.2̍W, 23–70 m, Sept. 6, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 ♀, 9.3 mm cl, allotype (MNHN­Hi 220) ; Sta. CP 1304, Île Nuku Hiva , 08 °54.4̍S, 140°13.9̍W, 50–58 m, Sept. 10, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 7.5 mm cl, paratype ( AMNH 17819 ) ; Sta. D 85, Île Fatu Hiva , 10°29.4̍S, 138°46.5̍W, 100 m, Jan. 29, 1991, coll. J. Poupin: 8 Ƌ, 5.2–10.0 mm cl, 2 ♀, 5.6–6.9 mm cl, 5 anterior half carapaces, 4.2–7.0 mm cl (MNHN­Hi 243), 1 ♀, 6.4 mm cl, paratype ( AMNH 17820 ) ; Sta. NH­VIII, Haul 5, Baie Hatuatua, Île Nuku Hiva, 08°51̍S, 140°00̍W, 26 fms (= 48 m), Sept. 18, 1967, coll. National Geographic Society –Smithsonian­ Bishop Museum Marquesas Expedition: 1 ♀, 5.2 mm cl, paratype ( USNM 260948 ) ; Sta. UP II, Haul 4, off west coast of Île Ua Pou, 40–45 fms (= 73–82 m), Sept. 23, 1967, coll. National Geographic Society – Smithsonian­Bishop Museum Marquesas Expedition: 1 Ƌ, 8.5 mm cl, paratype ( USNM 260949 ) ; Sta. FH I, Haul 1, west coast of Île Fatu Hiva, 10°27–30̍S, 138°40̍W, 41–43 fms (= 75–79 m), Sept. 25, 1967, coll. National Geographic Society – Smithsonian­Bishop Museum Marquesas Expedition: 1 anterior third ( USNM 260950 ) ; Sta. EO I, Haul 1, off northern coast of Île Eiao, 08°00̍S, 140°50̍W, 28–29 fms (= 51–53 m), Sept. 21, 1967, coll. National Geographic Society – Smithsonian­Bishop Museum Marquesas Expedition: 1 Ƌ, 7.9 mm cl, paratype ( USNM 260952 ) ; Sta. TH IX, Haul 1, off Hana Moe Noe, northwest coast of Île Tahuata, 09°54̍S, 139°07̍W, 37 fms (= 68 m), Oct. 1, 1967, coll. National Geographic Society – Smithsonian­Bishop Museum Marquesas Expedition: 1 Ƌ, 7.9 mm cl, 1 ♀, 8.4 mm cl, paratypes ( USNM 268578 ) ; Sta. DW 1143, Île Ua Pou , 09°20.9̍S, 140°02.7̍W, 18–55 m, Aug. 22, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 4.5 mm cl, paratype (MNHN­Hi 221) ; Sta. DW 1162, Île Nuku Hiva , 08°56.2̍S, 140°06.1̍W, 45–64 m, Aug. 24, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 anterior half, 4.7 mm cl (MNHN­Hi 222) ; Sta. DW 1185, Île Nuku Hiva , 08°48.9̍S, 140°03.4̍W, 31–33 m, Aug. 26, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 7.6 mm cl, 1 anterior half, 9.5 mm cl (MNHN­Hi 222 bis) ; Sta. DW 1180, Île Nuku Hiva , 08°46.2̍S, 140°04.6̍W, 80–82 m, Aug. 26, 1997, coll. MUSOR­ STOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 10.2 mm cl, paratype (MNHN­Hi 223) ; Sta. CP 1187, Île Nuku Hiva , 08°49.2̍S, 140°03.5̍W, 25–30 m, Aug. 26, 1997, coll. MUSOR­ STOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 3 Ƌ, 8.9– 10.8 mm cl, paratypes (MNHN­Hi 224) ; Sta. DW 1213, Île Hiva Oa , 09°50.3̍S, 139°03.2̍W, 18–20 m, Aug. 29, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 ♀, 5.3 mm cl (MNHN­Hi 225) ; Sta. DW 1214, Île Hiva Oa , 09°49.8̍S, 139°03.1̍W, 25–40 m, Aug. 29, 1997, coll. MUSOR­ STOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 anterior half, 7.1 mm cl (MNHN­Hi 226) ; Sta. DW 1217, Île Hiva Oa , 09°44.5̍S, 138°49.9̍W, 85–87 m, Aug. 30, 1997, coll. MUSOR­ STOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 5.1 mm cl, paratype (MNHN­Hi 227) ; Sta. DW 1241, Île Fatu Hiva , 10°27.8̍S, 138°40.6̍W, 85–130 m, Sept. 1, 1997, coll. MUSOR­ STOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 6.2 mm cl, paratype (MNHN­Hi 228) ; Sta. DW 1242, Île Fatu Hiva , 10°28.1̍S, 138°41.1̍W, 119–122 m, Sept. 1, 1997, coll. MUSOR­ STOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 7.2 mm cl, paratype (MNHN­Hi 229) ; Sta. DR 1245, Île Fatu Hiva , 10°29.2̍S, 138°36.2̍W, 85–130 m, Sept. 1, 1997, coll. MUSOR­ STOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 10.1 mm cl, 1 ♀, 9.4 mm cl, paratypes ( MNHN­ Hi 230) ; Sta. DW 1256, Île Ua Pou , 09°25.4̍S, 140°07.9̍W, 70–72 m, Sept. 3, 1997, MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 6 Ƌ, 5.2–8.7 mm cl, 2 ♀, 6.0– 7.4 mm cl, 1 posterior half of oviger ( MNHN­ Hi 231) ; Sta. DW 1260, Île Ua Pou , 09°25.4̍S, 140°07.3̍W, 49–100 m, Sept. 3, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 2 Ƌ, 8.4–8.9 mm cl, 1 ♀, 9.7 mm cl, paratypes (MNHN­Hi 232) ; Sta. DW 1266, Île Eiao , 07°57.3̍S, 140°42.6̍W, 84 m, Sept. 4, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 5.6 mm cl, paratype (MNHN­Hi 233) ; Sta. DW 1279, Île Eiao , 07°59.4̍S, 140°42.2̍W, 23–70 m, Sept. 6, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 4 ♀, 3.5–8.1 mm cl, paratypes (MNHN­Hi 234) ; Sta. DW 1283, Île Mutu One , Hatutaa, 07°53.8̍S, 140°34.5̍W, 55–56 m, Sept. 7, 1997, coll. MUSORSTOM 9 N/ O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 5.4 mm cl, paratype (MNHN­Hi 235) ; Sta. DW 1292, 08°54.1̍S, 139°37.8̍W, Ile Ua Huka , 95–100 m, Sept. 8, 1997, coll. MUSORSTOM 9 N/ O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 6.1 mm cl (MNHN­Hi 257) ; Sta. DW 1297, Île Ua Huka , 08°54.2̍S, 139°37.4̍W, 90–150 m, Sept. 8, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 1 Ƌ, 7.9 mm cl, paratype (MNHN­Hi 236), 1 ♀, 4.8 mm cl, paratype (MNHN­Hi 237) ; Sta. CP 1304, Île Nuku Hiva , 08°54.4̍S, 140°13.9̍W, 50–58 m, Sept. 10, 1997, coll. MUSORSTOM 9 N/O ‘‘ Alis’ ’ Campagne (P. Bouchet, B. Dayrat, B. Richer De Forges): 3 Ƌ, 6.3–8.0 mm cl, paratypes (MNHN­Hi 238) ; Sta. 24, Côte NW Baie Haahue, Île Ua Huka, 08°53.6̍S, 139°37̍W, 9–25 m, Oct. 1997, coll. R. Von Cosel, J. Tröndle´, and J. Tardy: 1 Ƌ, 4.3 mm cl, 4 ♀, 5.8–10.9 mm cl, 6 juveniles, 3.1–3.8 mm cl, 2 unsexable, unmeasurable specimens (MNHN­Hi 239) ; Sta. 29, Baie de Hane, Île Ua Huka, 08°55.7̍S, 139°32.0̍W, 7–11 m, Oct. 1997, coll. R. Von Cosel, J. Tröndle´, and J. Tardy: 2 ♀, 5.0– 5.1 mm cl (MNHN­Hi 240) ; Sta. 34, Baie Haavei , Pointe Tenoni , Île Teuaua, Île Ua Huka, 08°56.8̍S, 139°35.7̍W, 10–15 m, Oct. 1997, coll. R. Von Cosel, J. Tröndle´, and J. Tardy: 1 Ƌ, 6.3 mm cl, paratype (MNHN­Hi 241) ; Sta. D 77, Île Eiao , 07°56.7̍S, 139°30.8̍W, 54 m, Jan. 21, 1991, coll. J. Poupin: 1 Ƌ, 10.0 mm cl, paratype (MNHN­Hi 242) .

DIAGNOSIS: Carapace wider than long, covered with lightly setose grooves. Anterior margin with 8–10 spines on either side of ocular sinus. Setal field with narrow lateral elements and sinuous anterior margin. CG1 with separate posterior lateral elements; CG4 with two long lateral elements terminating above apex of CG6, occasionally with two short medial elements; CG5 broken into two short oblique elements; CG6 and CG7 separate; CG8 broken; CG11 present. Rostrum present, not reaching posterior margin of ocular plate. Ocular plate triangular. Distal peduncular segments dorsoventrally flattened and subtriangular in shape, tapering at tip, approximated along mesial margins, lateral margins convex except slightly concave at tip, mesial margins straight. Cornea at tip. Dactylus of pereopod II with heel produced and subquadrate. Dactylus of pereopod III with heel slightly produced and acute. Dactylus of pereopod IV evenly sinuous from base to tip. Telson of male spatulate, tip rounded, dorsal surface inflated medially with strong medial ridge lined with short thick setae; lateral margins decalcified. Telson of female ovate.

DESCRIPTION: Carapace (fig. 92A) slightly wider than long. Anterior margin concave on either side of ocular sinus, becoming convex laterally, with 8–10 large spines along length. Rostrum as small acute tooth, not reaching proximal margin of ocular plate. Ocular sinus smoothly concave and unarmed. Frontal region smooth; setal field broad posteriorly, narrowing anteriorly, with narrow anterior lateral elements and sinuous anterior margin; posterior lateral elements not reaching to posterior lateral elements of CG1. CG1 parallel to anterior margin of carapace, sinuous, slightly crenulate, divided into medial sinuous fragment and curved, posteriorly displaced lateral elements. Mesogastric region smooth; CG2 absent; CG3 broken into four to six short elements; CG4 with two long lateral elements terminating above apex of CG6, occasionally with two short medial elements. Hepatic region smooth, with long setose groove at median of lateral margin. Epibranchial region generally triangular, smooth. Metagastric region smooth; CG5 broken into two short oblique elements. CG6 strongly crenulate, strongly anteriorly concave medially and sloping out to anteriorly convex lateral thirds. CG7 present as two long oblique elements and separate from CG6. Cardiac region smooth; CG8 with two to four median elements and two slightly longer lateral elements. CG9 present as two short, widely separated elements. CG10 present as two straight lateral fragments, with gap between fragments greater than length of single fragment. CG11 present as two or three irregularly spaced short elements. Branchial region with numerous short, transverse rows of setae. Posterior margin deeply and evenly convex, with submarginal groove reaching approximately three­fourths up either side of posterior concavity. Branchiostegite with short anterior submarginal spine; anterior region with scattered short, transverse lines ventral to linea anomurica; with many short rows of setae and sparsely covered with long plumose setae ventrally; posterior region membranous, with numerous irregular fragments and sparsely covered with long plumose setae.

Ocular plate (fig. 92B) subquadrate, with shallow median indentation; median peduncular segments reduced to small rounded calcified area on either side of ocular plate. Distal peduncular segments elongate, with proximally convex and distally concave lateral margins, tapering to produced distal cornea; mesial margins approximated along entire length; mesial and ventral margins of segment with sparse row of long plumose setae; few small tufts of plumose setae on proximal quarter of segment.

Antennule (fig. 92C) with segment III subcylindrical; with plumose setae on dorsal and ventral margins; dorsal exopodal flagellum with 110–118 articles, long plumose setae on dorsal and ventral margins; ventral endopodal flagellum short with two articles and plumose setae on dorsal and ventral margins. Segment II medially inflated in dorsal view, with plumose setae on dorsal and ventral margins and scattered on ventrolateral third of surface. Segment I wider than long, unarmed; dorsal third of lateral surface rugose with long plumose setae; long plumose setae on dorsal and ventral margins.

Antenna (fig. 92D) with segment V approximately two times longer than wide, with long plumose setae on dorsal margins; flagellum with seven articles, long plumose setae on dorsal, ventral, and distal margins. Segment IV expanded distally, with long plumose setae on dorsal, ventral, and distal margins and row of setae on dorsolateral margin. Segment III with long plumose setae on dorsal and ventral margins. Segment II short, widening distally, with plumose setae on margins; antennal acicle long, thin, truncate distally, slightly exceeding distal margin of segment IV, with long plumose setae on dorsal margin. Segment I rounded proximally, flattened ventrolaterally, with long plumose setae on margins; lateral surface with acute spine subdorsally, with low semicircular, dorsolateral lobe ventrodistal to spine; segment with ventromesial antennal gland pore.

Mandible (fig. 92E) incisor process without teeth; cutting edge with one tooth. Palp three­segmented, with plumose setae on margins and long, thick, simple setae arising from bend in second segment.

Maxillule (fig. 92F) distal endite proximally narrow, widening to inflated distal end, with thick simple setae on distal margin; plumose setae on dorsal margin. Proximal endite with thick simple setae on distal margin. Endopodal external lobe truncate distally and curled under; internal lobe reduced with three thick setae at distolateral margin.

Maxilla (fig. 92G) exopod evenly rounded, with plumose setae along distal margin. Scaphognathite bluntly angled on posterior lobe, with plumose setae.

Maxilliped I (fig. 92H) epipod with plumose setae on distal margin and on distolateral surface. Endite tapered distally and subequal to first segment of exopod. Exopod with two segments; proximal segment narrow, margins parallel, with plumose setae; distal segment spatulate, approximately as long as wide, broadest medially, margins with long plumose setae. Endopod flattened and elongate, reaching to distal end of proximal exopodal segment, with plumose setae on margins.

Maxilliped II (fig. 92I) dactylus evenly rounded, length slightly greater than width, with thick simple setae distally. Propodus two times wider than long, with plumose setae on dorsal margin and long simple setae on distal margin. Carpus not strongly produced dorsodistally, approximately three times longer than wide, with long simple setae on dorsal margin. Merus approximately three times longer than wide, margins parallel, with simple setae on ventrolateral margin and scattered on surface, plumose setae on dorsolateral margin. Basis­ischium incompletely fused, with plumose setae on margins. Exopod one­fourth longer than merus, flagellum with one elongate article.

Maxilliped III (fig. 92J) dactylus evenly rounded; with long plumose setae on margins and lateral surface. Propodus with longitudinal median row of plumose setae on lateral surface; margins with plumose setae. Carpus slightly produced onto propodus; lateral surface with row of plumose setae ventromedially; plumose setae on margins. Merus unarmed, with plumose setae on margins and scattered on surface. Basis­ischium incompletely fused, with faint crista dentata of few small low teeth. Exopod two­segmented: proximal segment small; distal segment styliform, tapering, approximately one­third length of merus, with plumose setae on margins; without flagellum.

Pereopod I (fig. 93A) dactylus curved and tapering; lateral and mesial surfaces smooth; dorsal margin with long plumose and short simple setae; ventral margin with short simple setae. Propodal lateral surface with numerous short, transverse rows of setose rugae; dorsal margin unarmed; ventral margin distally produced into acute spine; cutting edge lacking teeth, lined with long plumose setae; dorsal margin with long plumose setae, ventral margin with short simple setae. Carpus with dorsodistal angle produced into small spine, dorsal margin smooth; dorsal and distal margins with long plumose setae; lateral surface with small distal rugose area, few transverse setose ridges on distal half of surface; mesial surface smooth, with median row of long plumose setae, margins with long plumose setae. Merus unarmed; lateral surface with scattered transverse rows of long plumose setae, margins with long plumose setae; mesial surface with few short rows of setae; proximal fourth of mesial surface with decalcified window. Basis­ischium incompletely fused, unarmed. Coxa unarmed.

Pereopod II (fig. 93B) with dactylus smooth; with base to heel slightly concave, heel smoothly rounded and slightly produced, heel to tip with acute, narrow indent, tip acute, tip to base broadly convex; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, several widely spaced submarginal tufts of short setae dorsodistally; mesial surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae, with patch of long plumose setae at base. Propodus with dorsal surface smooth, ventral margin inflat­ ed and rounded; oblique row of long plumose setae on distal margin of lateral surface; distal and ventral margin with long plumose setae; dorsolateral surface as narrow, oblique, flattened shelf, with short setae on dorsal margin and long plumose setae on ventral margin; mesial surface with elevated, curved, setose ridge from ventral junction with dactylus almost to ventral proximal junction with carpus; decalcified region just distal to junction with carpus. Carpus slightly produced dorsodistally; lateral surface nearly smooth, with irregular, interrupted row of rugae and submarginal elevated ridge ventrally, rugae and ridge with long plumose setae; margins with long plumose setae; mesial surface smooth with long plumose setae in scattered patches on surface and on margins. Merus lateral surface with large decalcified area in median, few scattered setae on surface and margins and large patch of long simple setae at distolateral margin; mesial surface nearly smooth, with many median setae, with decalcified area on proximal fourth near junction with basis­ischium. Basis­ischium incompletely fused and unarmed. Coxa with one small tubercle on anterior margin.

Pereopod III (fig. 93C) dactylus with base to heel concave, heel broadly rounded and slightly produced, heel to tip with broad, evenly rounded indent, tip acute, tip to base smoothly convex to straight; lateral surface smooth, with several small tufts of short setae in generally straight line across medioproximal surface, dorsodistal margin with tufts of short setae; ventromesial margin with long plumose setae, dorsal margin with short simple and plumose setae; mesial surface smooth, with plumose setae proximally at junction with propodus. Propodus not inflat­ ed dorsoventrally; lateral surface smooth, with long plumose setae distally, with simple setae on dorsal margins, and long plumose setae on ventral margin; dorsolateral surface narrow, oblique, flattened; mesial surface with scattered long setae on and near distal margin, with decalcified window near junction with carpus. Carpus produced dorsodistally, exceeding proximal margin of propodus by approximately one­fourth length of propodus, pointed but not acute; dorsolateral margin unarmed; lateral surface slightly rugose dorsodistally, with mat of short setae and two longer rows of setae ventrally; mesial surface smooth, with long plumose setae on margins and in transverse row on surface. Merus smooth, with large decalcified area near median of lateral surface; dorsal and ventral margins unarmed, with long plumose setae; distolateral margin with long plumose setae; mesial surface smooth, with decalcified window at junction with basis­ischium. Basis­ischium incompletely fused and unarmed. Coxa unarmed. Female with large gonopore on anterior mesial surface of coxa, surrounded with short plumose setae; male without pore.

Pereopod IV (fig. 93D) dactylus with base to tip convex to concave, tip acute, tip to base straight distally, becoming convex proximally; lateral surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae; mesial surface with dorsal decalcified region, demarcated ventrally by longitudinal elevated ridge with row of short setae; with setose punctae ventral to decalcified window. Propodus expanded dorsally and ventrally; ventral expansion not exceeding ventral margin of dactylus, margin with long plumose setae; dorsal expansion with row of long plumose setae medially and mat of short simple setae; lateral and mesial surfaces smooth, mesial surface with large decalcified area. Carpus not produced dorsodistally; lateral and mesial surfaces smooth; dorsal margin with short simple and long plumose setae; ventral margin with short simple setae and small mat of short simple setae dorsally. Merus with scattered, short, transverse rows of setae on lateral surface, dorsal and ventrodistal margins with long plumose setae; mesial surface with large decalcified window proximoventrally. Basisischium incompletely fused and unarmed. Coxa unarmed.

Abdomen (fig. 93E) with somite I approximately as long as wide, widest posteriorly; dorsal surface with anterior margin straight; posterior margin concave, with elevated submarginal row of short setae; with small transverse, decalcified windows laterad of segment median. Somite II dorsal surface with submarginal transverse ridge anteriorly; with small transverse, decalcified windows laterad of segment median just anterior to submarginal ridge; with tuft of setae at posterolateral angle, extending onto pleura posteromesially; posterior margin with indistinct punctate submarginal groove laterally; pleura expanded and directed slightly anteriorly; lateral margins rounded, anterior and lateral margins with long plumose setae, posterior margin with short setae. Somite III similar to somite II, but narrower, shorter, and lacking anterior submarginal ridge; small tuft of short thick setae on posterolateral angle; pleura thinner and shorter than on somite II, directed anterolaterally, with setae as in somite II; anterolateral angle acute; dorsal surface obliquely flattened anterolaterally. Somite IV similar to somite III, but thinner and shorter; dorsal surface with thick setae posterolaterally; pleura thinner and shorter than on somite III, directed posterolaterally; dorsal surface obliquely flattened anterolaterally; margin with long plumose setae. Somite V subequal to somite IV; lateral margins with plumose setae; pleura absent. Somite VI subequal to somite V in width but longer; dorsal surface with short transverse rows of setae laterad of midline anteriorly; pleura absent.

Females with uniramous, paired pleopods on somites II–V; males lacking pleopods.

Telson of male (fig. 93F) spatulate, with length greater than width, rounded distally; weakly calcified at margins of large calcified median plate; median longitudinal groove short, extending one­fourth length of telson; thick elevated ridge continuing from end of groove to distal end of telson, lined with dense row of thick simple setae. Telson of female (fig. 93G) ovate, longer than wide, dorsal surface smooth, with median longitudinal groove reaching almost to distal margin; with row of setose punctae lateral to midline along whole length of median groove except proximal fourth; margins with long plumose setae.

DISTRIBUTION: Known only from the Marquises Islands, in 7–130 m depth.

MAXIMUM SIZE: Males: 10.8 mm cl; females: 10.9 mm cl.

TYPE SPECIMENS: USNM 268577 (holotype), MNHN­Hi 220 (allotype), AMNH 17819 (paratype), AMNH 17820 (paratype), USNM 260948 (paratype), USNM 260949 (paratype), USNM 260952 (paratype), USNM 268578 (2 paratypes), MNHN­Hi 221 (paratype), MNHN­Hi 223 (paratype), MNHN­Hi 224 (3 paratypes), MNHN­Hi 227 (paratype), MNHN­Hi 228 (paratype), MNHN­Hi 229 (paratype), MNHN­Hi 230 (2 paratypes), MNHN­Hi 232 (3 paratypes), MNHN­Hi 233 (paratype), MNHN­Hi 234 (4 paratypes), MNHN­Hi 235 (paratype), MNHN­Hi 236 (paratype), MNHN­Hi 237 (paratype), MNHN­Hi 238 (3 paratypes), MNHN­Hi 241 (paratype), MNHN­Hi 242 (paratype).

TYPE LOCALITY: Haava Straits between Île Tahuata and Île Hiva Oa, Marquises Islands, 09°52̍S, 139°04̍W, 73 m.

REMARKS: Juvenile specimens are virtually lacking in pigment and appear almost white. Adults are a uniform tan, with reddish­brown setae. Larger specimens show a markedly increased reddish tone on the carapace (especially the anterior region), ocular peduncles, antennae, and antennules (from preserved material and a color transparency made of a live specimen [MNHN­Hi 223]; see also Poupin, 1996b).

Little is known about the biology of this species. Ovigerous females, unfortunately, are only known from the posterior portion of a single specimen. Additionally, the morphology of the spermatozoa of this species has been studied by Tudge et al. (1999), based on one of the paratypes (MNHN­Hi 236).

Albunea marquisiana is most similar to A. holthuisi and A. groeningi . All three species share the distinctive thick median row of setae on the telson of the male, as well as a general similarity in the shape of the pereopodal dactyli and can be collectively considered to form the ‘‘ holthuisi ­group’’ of species. They can easily be separated by the number of elements of CG8 (one long median element in A. holthuisi , three or four short elements in A. marquisiana and A. groeningi ), and CG11 (absent in A. groeningi , one in A. holthuisi , two or three in A. marquisiana ), the relative thickness of the proximal blade of pereopod III (thicker in A. marquisiana and A. groeningi ), and the distal margins of both the male and female telsons (truncate in A. holthuisi , smoothly rounded in A. marquisiana , indented in A. groeningi ).

Albunea holthuisi Boyko and Harvey, 1999 Figures 94 , 95

? Albunea symnista [sic]: Ward, 1942: 52 (list), 63

(not Albunea symmysta ( Linnaeus, 1758)) .? Albunea steinitzi: Thomassin, 1969: 143–146 , pl.

3, figs. 1–8, text­figs. 3c, 4 (not Albunea stein­

itzi Holthuis, 1958). Albunea holthuisi Boyko and Harvey, 1999: 386–

391, 400 (list), 401 (key), figs. 6, 7*.

MATERIAL EXAMINED: Zanzibar: Sta. 651, dredged grass and shell, 1.5 mi west­southwest of Ras Nungwa , 8 fms (= 14.6 m), Feb. 20, 1957, coll. A. J. Ostheimer III: 1 Ƌ, 7.7 mm cl, paratype ( ANSP CA4644 ) ; Sta. 651, fine grass and shell, 1.5 mi west­southwest of Ras Nungwa , 8 fms (= 14.6 m), Feb. 20, 1957, coll. A. J. Ostheimer III: 1 ♀, 6.7 mm cl, paratype ( ANSP CA4645 ) .

Madagascar: Environs de Nosy Be , Côte southwest, 13°37.7̍S, 47°49.6̍E, 25 m, coll. unknown: 1 Ƌ, 8.1 mm cl, holotype (MNHN­Hi 202) ; Andilana , Nosy Be, Côte southwest, Sept. 1959, coll. A. Crosnier: 1 ♀, 8.1 mm, allotype (MNHN­Hi 203) ; voisinage de Nosy Be, Côte northwest, 13°38.3̍S, 42°49.6̍E, 34 m, coll. A. Crosnier: 1 Ƌ, 8.0 mm cl, paratype (MNHN­Hi 204) ; Sakatia , Envoi II, 1921, coll. G. Petit: 1 Ƌ, 6.7 mm cl (MNHN­Hi 19) .

Seychelles: Sta. 5, 33 m, Sept. 4, 1980, coll. ORSTOM­Reves 2: 1 Ƌ, 9.2 mm cl (MNHN­Hi 249) ; Mahe´, July–Aug. 1972, coll. Mission Zoologique MRAC­ULB: 1 ♀, 5.7 mm cl ( MRAC 57.459 ) .

Indonesia: Sta. 522, west side of Samberbaban Bay, Japen Island, Irian Jaya, Feb. 14, 1956, coll. C. T. Abbot on ‘‘ Gloria Maris’ ’: 1 Ƌ, 11.3 mm cl, paratype ( RMNH 23703 ) ; Corindon II, Sta. B255, Makassar, 01°56.5̍S, 119°17.3̍E, 13 m, Nov. 6, 1980, coll. R/V ‘‘Coriolis’’: 1 ♀, 7.1 mm cl, paratype (MNHN­Hi 205); Corindon II, Sta. B256, Makassar, 01°56.5̍S, 119°17.2̍E, 24 m, Nov. 6, 1980, coll. R/V ‘‘Coriolis’’: 1 Ƌ, 4.9 mm cl, 1 ♀, 5.4 mm cl (MNHN­Hi 206).

Malaysia: Juara Bay, Pulau Tioman, Pahang, May 26, 1985, coll. P.K.L. Ng: 1 Ƌ, 7.5 mm cl ( ZRC 1989.3827).

Australia: Queensland: Little Trunk Reef, 18°20̍S, 146°46̍E, 9.1–12.2 m, Nov. 5, 1990, coll. K. Lamprell: 1 Ƌ, 9.6 mm cl ( QM W24961).

DIAGNOSIS: Carapace slightly longer than wide, covered with lightly setose grooves. Anterior margin with 8–11 spines on either side of ocular sinus. Setal field with narrow lateral elements and concave anterior margin. CG1 with separate posterior lateral elements; CG4 with short element on either side of median with missing elements at midline and between median and laterals; CG5 present only as short lateral elements; CG6 and CG7 separate; CG8 complete; CG11 present. Rostrum present, not reaching posterior margin of ocular plate. Ocular plate triangular. Distal peduncular segments dorsoventrally flattened and oblong in shape, tapering at tip, approximated along distal two­thirds of mesial margins, lateral margins convex except slightly concave at tip, mesial margins convex. Cornea at tip. Dactylus of pereopod II with heel slightly produced, low and rounded. Dactylus of pereopod III with heel slightly projecting, acute. Dactylus of pereopod IV sinuous from base to tip, with slight indent. Telson of male spatulate, tip broadly truncate, dorsal surface with elevated, median, longitudinal ridge bearing short thick setae proximally and long thick setae distally. Telson of female flattened and spatulate, longitudinal row of short, thin setae medially.

DESCRIPTION: Carapace (fig. 94A) slightly wider than long. Anterior margin concave on either side of ocular sinus, becoming convex laterally, 8–11 large spines on concave region, ventral row of long plumose setae submarginally. Rostrum as small acute tooth, not reaching to proximal margin of ocular plate. Ocular sinus smoothly concave and unarmed. Frontal region smooth; setal field broad posteriorly, narrowing anteriorly, with narrow lateral elements and concave anterior margin. CG1 parallel to anterior margin of carapace, sinuous, slightly crenulate, divided into medial fragment and curved lateral elements that are displaced posteriorly. Mesogastric region smooth, CG2 short; CG3 broken into six short elements approximately equally spaced between posterior elements of CG1; CG4 fragmented into four elements with gap at midline and between median and lateral elements. Hepatic region smooth, with short setose groove at median of lateral margin. Epibranchial region generally triangular, smooth, posterolateral margin with short row of setae. Metagastric region smooth; CG5 present only as short lateral elements directly posterior to median elements of CG4; CG6 slightly crenulate, strongly concave medially and sloping out to convex lateral thirds; CG7 transverse and separate from CG6. Cardiac region smooth; CG8 uninterrupted; CG9 present only as lateral short lines; CG10 present in two fragments, separated by length of sin­ gle fragment; CG11 present. Branchial region with numerous short, transverse rows of setae. Posterior margin deeply and evenly convex, with submarginal groove interrupted medially. Branchiostegite with short anterior submarginal spine, anterior region with scattered short, transverse lines ventral to linea anomurica, with many short rows of setae and covered with long plumose setae ven­ trally, posterior region membranous, with numerous irregular fragments, and covered with long plumose setae.

Ocular plate (fig. 94B) triangular, with shallow median indentation; median peduncular segments present as small, ovate calcified areas lateral to ocular plate. Distal peduncular segments elongate, with distally convex lateral margins, tapering to rounded distolateral cornea, mesial margins approximated along entire length, mesial and ventral margins of segment with sparse row of long plumose setae, tuft of plumose setae at proximal lateral ventral angle, ventral surface with oblique row of long plumose setae from proximal lateral angle almost to distal mesial margin.

Antennule (fig. 94C) with segment III narrow proximally, expanding distally to twice proximal width; plumose setae on dorsal and ventral margins, dorsal exopodal flagellum with 76–104 articles, long plumose setae on dorsal and ventral margins, ventral endopodal flagellum short, with two or three articles, plumose setae on dorsal and ventral margins. Segment II medially inflated in dorsal view, plumose setae on dorsal and ventral margins and scattered on lateral surface. Segment I wider than long, unarmed, lateral surface with long plumose setae dorsally and on dorsal and ventral margins.

Antenna (fig. 94D) with segment V approximately three times longer than wide, long plumose setae on dorsal and ventral margins, flagellum with seven articles, long plumose setae on dorsal, ventral, and distal margins. Segment IV expanded distally, long plumose setae on dorsal, ventral, and distal margins and simple setae on dorsolateral margin. Segment III with long plumose setae on ventral margin. Segment II short, widening distally, plumose setae on margins, antennal acicle long, thin, exceeding base of segment V by approximately one­half length of segment V, long plumose setae on dorsal margin. Segment I rounded proximally, flattened ventromesially, long plumose setae on margins; lateral surface with acute spine dorsally, with low, semicircular dorsolateral lobe ventrodistal to spine.

Mandible (fig. 94E) incisor process with one tooth; cutting edge with one tooth. Palp three­segmented, with plumose setae on mar­ gins and long, thick, simple setae arising from bend in second segment.

Maxillule (fig. 94F) distal endite proximally narrow, widening to inflated distal end, with thick simple setae on distal margin. Proximal endite with thick simple setae on distal margin. Endopodal external lobe truncate distally and curled under, notched proximally; internal lobe reduced with two thick setae at distolateral margin.

Maxilla (fig. 94G) exopod evenly rounded, with plumose setae along distal margin. Scaphognathite bluntly angled on posterior lobe, with plumose setae.

Maxilliped I (fig. 94H) with epipod with plumose setae on distal margin and on distolateral surface. Endite tapered distally and subequal to first segment of exopod. Exopod with two segments; proximal segment narrow, margins parallel, with plumose setae; distal segment spatulate, slightly longer than wide, broadest medially, margins with long plumose setae. Endopod flattened and elongate, reaching to distal end of proximal exopodal segment, with plumose setae on margins.

Maxilliped II (fig. 94I) dactylus evenly rounded, length equal to width, with thick simple setae distally. Propodus 1.5 times wider than long, with plumose setae on dorsal margin and long simple setae on distal margin. Carpus not strongly produced dorsodistally, approximately two times longer than wide, with long simple setae on dorsal margin. Merus approximately three times longer than wide, margins parallel, with simple setae on ventrolateral margin and plumose setae on dorsolateral margin. Basis­ischium incompletely fused, with plumose setae on margins. Exopod one­fourth longer than merus, flagellum with one article.

Maxilliped III (fig. 94J) dactylus rounded at tip, long plumose setae on margins and lateral surface. Propodus with longitudinal median row of plumose setae on lateral surface, margins with plumose setae. Carpus slightly produced onto propodus, lateral surface with row of plumose setae ventromedially; plumose setae on margins. Merus unarmed, plumose setae on margins. Basis incompletely fused with ischium; weak crista dentata of two or three teeth. Exopod twosegmented, proximal segment small, distal segment styliform, tapering, approximately one­third length of merus, plumose setae on margins; flagellum absent.

Pereopod I (fig. 95A) dactylus curved and tapering; lateral and mesial surfaces smooth; dorsal margin with long plumose and short simple setae, short simple setae on ventral margin. Propodal lateral surface with numerous short, transverse rows of setose rugae; dorsal margin unarmed; ventral margin dis­ tally produced into acute spine; cutting edge lacking teeth, lined with long plumose setae; lateral, mesial, and ventral margins with long setae. Carpus with dorsodistal angle produced into small corneous spine; dorsal and distal margins with long plumose setae; lateral surface with distal rugose area, few transverse setose ridges on distal two­thirds of surface; mesial surface smooth, with scattered rows of long plumose setae, margins with long plumose setae. Merus unarmed; lateral surface with scattered transverse rows of long plumose setae, margins with long plumose setae; mesial surface with few short rows of setae. Basis­ischium incompletely fused, unarmed. Coxa unarmed.

Pereopod II (fig. 95B) dactylus smooth; base to heel concave, heel with smoothly rounded low spur, heel to tip acutely indented and narrow, tip acute, tip to base broadly convex; lateral surface smooth, several small tufts of short setae in generally straight line across medioproximal surface, several widely spaced submarginal tufts of short setae dorsodistally; mesial surface smooth, ventral margin with long plumose setae, dorsal margin with short plumose setae, patch of long plumose setae at base (not illustrated). Propodal dorsal surface smooth, ventral margin inflated and rounded; oblique row of long plumose setae on distal margin of lateral surface; distal and ventral margins with long plumose setae; dorsolateral surface as narrow, oblique, flattened shelf, short setae on dorsal margin and long plumose setae on ventral margin; mesial surface with elevated, curved, setose ridge from ventral junction with dactylus almost to ventral proximal junction with carpus. Carpus slightly produced, gently rounded; lateral surface nearly smooth, with irregular, broken row of rugae and submarginal elevated ridge ventrally, rugae and ridge with long plumose setae, margins with long plumose setae; mesial surface smooth, long plumose setae on margins and in scattered patches on surface. Merus with medial decalcified area on lateral surface, long plumose setae on lateral margins; mesial surface nearly smooth, with few setae. Basis­ischium incompletely fused and unarmed. Coxa with one small spine on anterior margin.

Pereopod III (fig. 95C) dactylus with base to heel concave, heel produced in short, acute spur, heel to indent nearly straight, indent broadly concave, tip acute, tip to base smoothly convex to straight; lateral surface smooth, dorsodistal margin with tufts of short setae, ventromesial margin with long plumose setae, dorsal margin with short simple and plumose setae; mesial surface smooth, plumose setae proximally at junction with propodus. Propodus weakly inflated; lateral surface smooth, long plumose setae distally, simple setae on margins, long plumose setae on ventral margin, dorsolateral surface narrow, oblique, flattened; mesial surface with scattered long setae on and near distal margin. Carpus produced dorsodistally, exceeding proximal margin of propodus by approximately one­third length of propodus, broadly rounded, dorsolateral margin unarmed; lateral surface slightly rugose dorsodistally, many short and two longer rows of setae ventrally; mesial surface smooth, long plumose setae on margins and scattered on surface. Merus smooth, dorsal and ventral margins unarmed, long plumose setae, distolateral margin with long plumose setae; lateral surface with decalcified area anteriorly; mesial surface smooth. Basis­ischium incompletely fused and unarmed. Coxa with one small spine on anterior margin. Female with large gonopore on median mesial surface of coxa, surrounded with short plumose setae; male without pore.

Pereopod IV (fig. 95D) dactylus with base to tip convex to straight, tip acute, tip to base convex distally, becoming broadly concave proximally; lateral surface smooth, ventral margin with long plumose setae, dorsal margin with short simple setae; mesial surface with median decalcified area, demarcated ventrally by longitudinal elevated ridge with row of long plumose setae, setose punctae ventral to decalcified window. Propodus expanded dorsally and ventrally, ventral expansion exceeds ventral margin of dactylus, margin with long plumose setae, dorsal expansion with row of long plumose setae medially; lateral and mesial surfaces smooth. Carpus not produced dorsodistally; lateral and mesial surfaces smooth, dorsal margin with short simple and long plumose setae, ventral margin with short plumose setae. Merus with scattered short, transverse rows of setae on lateral surface, dorsal and ventrodistal margins with long plumose setae, slightly rugose ventrodistally, with short setae; mesial surface with large decalcified window proximoventrally. Basis­ischium incompletely fused and unarmed. Coxa unarmed.

Abdomen (fig. 95E) with somite I approximately as wide as long, widest posteriorly; dorsal surface with anterior margin concave, posterior margin concave, with submarginal row of short setae, small transverse decalcified submedial windows. Somite II dorsal surface with submarginal transverse ridge anteriorly, tuft of setae at posterolateral angle, extending onto pleura posteromesially, posterior margin with indistinct punctate submarginal groove laterally; pleura expanded and directed slightly anteriorly, margins finely toothed, lateral margins rounded, anterior and lateral margins with long plumose setae, posterior margin with short setae. Somite III similar to somite II, but narrower, shorter, and lacking anterior submarginal ridge, small tuft of short thick setae on posterolateral angle; pleura thinner and shorter than on somite II, directed anterolaterally, with setae as in somite II, anterolateral angle acute, dorsal surface obliquely flattened anterolaterally. Somite IV similar to somite III, but thinner and shorter; dorsal surface with thick setae posterolaterally; pleura thinner and shorter than on somite III, directed laterally, dorsal surface obliquely flattened anterolaterally, margins with long plumose setae. Somite V wider than somite IV, lateral margins with plumose setae; pleura absent. Somite VI subequal to somite V in width but longer, dorsal surface with short oblique rows of setae laterad of midline anteriorly, lateral margins with long plumose setae; pleura absent.

Females with uniramous, paired pleopods on somites II–V, males lacking pleopods.

Telson of male (fig. 95F, G) spatulate, truncate posteriorly, weakly calcified, except for large triangular anterior plate, margins with long plumose setae; median longitudinal groove very short, restricted to anterior of calcified plate, calcified plate with thick elevated medial ridge (fig. 95G) covered with short thick simple setae, small tuft of setae at anterolateral margin. Telson of female (fig. 95H) ovate, longer than wide, slightly trun­ cate posteriorly, dorsal surface smooth, with median longitudinal groove anteriorly, row of setose punctae lateral to midline from median of longitudinal groove almost to posterior margin; margins with long plumose setae.

DISTRIBUTION: Madagascar, Zanzibar, and the Seychelles; also from Indonesia and Queensland, Australia, in 9.1–34.0 m depth.

MAXIMUM SIZE: Males: 11.3 mm cl; females 8.1 mm cl.

TYPE SPECIMENS: MNHN­Hi 202 (holotype), MNHN­Hi 203 (allotype), MNHN­Hi 204 (paratype), MNHN­Hi 205 (paratype), ANSP CA4644 (paratype), ANSP CA4645 (paratype), RMNH 23703 (paratype).

TYPE LOCALITY: Environs de Nosy Be, Côte southwest, Madagascar, 13°37.7̍S, 47°49.6̍E, 25 m.

REMARKS: As with all older records of Albunea species lacking descriptive information or illustrations, it is difficult to be certain what species Ward (1942) was dealing with from Mauritius. I have examined no specimens from that locality, and the only species known from nearby Réunion is A. speciosa , which no doubt Ward (1942) would have recognized as quite different from A. symmysta . Albunea holthuisi is the best candidate for Ward’s (1942) record, as it superficially resembles A. symmysta and is known from nearby Madagascar and the Seychelles. The other taxa known from the vicinity of Mauritius ( A. elioti and A. microps ) are unlikely to have been confused with A. symmysta by an experienced carcinologist such as Ward.

As noted by Boyko and Harvey (1999), the record of ‘‘ Albunea steinitzi ’’ from Madagascar given by Thomassin (1969) is not that species. This record is tentatively placed in synonymy with A. holthuisi , which is also known from Madagascar, even though certain morphological details shown in Thomassin’s (1969) drawings do not fit perfectly with this species. Thomassin’s (1969) drawing of the habitus has a produced distal margin of the (presumably male) telson, while the indent of the dactylus of pereopod II is too broad. These may, however, be due to the relatively poor quality of Thomassin’s (1969) drawings (see under A. speciosa ). Thomassin’s (1969) record may represent an undescribed species, but as his material is un­ available for examination (see under A. speciosa ), no definite conclusions can be made at this time.

Although Boyko and Harvey (1999) indicated that this species is most similar to A. steinitzi , that relationship was based on the species described at that time. In fact, A. holthuisi typifies its own group of Albunea species , and it is actually the sister taxon to A. groeningi . It is also closely related to A. marquisiana .

Figures

Fig. 87. Albunea carabus (Linnaeus, 1758): A, Ƌ, 21.0 mm cl, USNM 258373; B–J, Ƌ, 19.1 mm cl, USNM 258373. A. Carapace, branchiostegite, and ocular peduncles, dorsal view. B. Ocular peduncles, dorsal view. C. Left antennule, lateral view. D. Left antenna, lateral view. E. Left mandible, mesial view. F. Left maxillule, lateral view. G. Left maxilla, lateral view. H. Left maxilliped I, lateral view. I. Left maxilliped II, lateral view. J. Left maxilliped III, lateral view. Scale = 2.2 mm (B), 3.3 mm (E, F), 4.4 mm (I), 6.7 mm (C, D, G, H, J), and 8.5 mm (A).

Fig. 88. Albunea carabus (Linnaeus, 1758): A, B, F, Ƌ, 19.1 mm cl, USNM 258373; C, D, Ƌ, 21.0 mm cl, USNM 258373; E, Ƌ, 19.1 mm cl, USNM 258373; G, ♀, 23.5 mm cl, RMNH 28911. A. Left pereopod I, lateral view. B. Right pereopod II, lateral view. C. Left pereopod III, lateral view. D. Left pereopod IV, lateral view. E. Abdominal somites I–VI, dorsal view. F. Telson of Ƌ, dorsal view. G. Telson of ♀, dorsal view. Scale = 3.3 mm (F, G), 6.7 mm (A, B), and 8.4 mm (C–E).

Fig. 89. Albunea asymmetrica (Müller, 1979): A. Anterolateral left quarter of carapace, 10.5 × 13.6 mm (l × w), HNHM MGy­2–1, holotype. B. Posterolateral right quarter of carapace, 13.8 × 12.2 mm (l × w), HNHM, paratype. C. Right half of carapace, 20.2 × 10.1 mm (l × w), HNHM MGy­2, paratype. D. Anterior section of right branchiostegite, 12.5 × 9.5 mm (l × w), HNHM. Scale = 6.7 mm (B–D) and 7.2 mm (A).

Fig. 90. Albunea danai Boyko, 1999: A, Ƌ, 16.7 mm cl, QM W23105, holotype; B–F, H–J, oviger, 10.8 mm cl, WAM 10422, paratype; G, Ƌ 11.6 mm cl, BPBM S5343, paratype. A. Carapace, branchiostegite, and ocular peduncles, dorsal view. B. Ocular peduncles, dorsal view. C. Left antennule, lateral view. D. Right antenna, lateral view. E. Left mandible, mesial view. F. Left maxillule, lateral view. G. Left maxilla, lateral view. H. Left maxilliped I, lateral view. I. Left maxilliped II, lateral view. J. Left maxilliped III, lateral view. Scale = 1.2 mm (C, D, G, H, J), 1.6 mm (B, E, F), 2.2 mm (I), and 6.7 mm (A).

Fig. 91. Albunea danai Boyko, 1999: A, F, Ƌ, 16.7 mm cl, QM W23105, holotype; B–E, Ƌ, 11.6 mm cl, BPBM S5343, paratype; G, ♀, 16.8 mm cl, BPBM S11782, allotype. A. Left pereopod I, lateral view. B. Right pereopod II, lateral view. C. Right pereopod III, lateral view. D. Right pereopod IV, lateral view. E. Abdominal somites I–VI, dorsal view. F. Telson of Ƌ, dorsal view. G. Telson of ♀, dorsal view. Scale = 3.0 mm (F, G) and 4.4 mm (A–E).

Fig. 92. Albunea marquisiana Boyko, 2000: A, Ƌ, 10.2 mm cl, USNM 268577, holotype; B–J, Ƌ, 7.9 mm cl, USNM 268578, paratype. A. Carapace, branchiostegite, and ocular peduncles, dorsal view. B. Ocular peduncles, dorsal view. C. Left antennule, lateral view. D. Left antenna, lateral view. E. Left mandible, mesial view. F. Left maxillule, lateral view. G. Left maxilla, lateral view. H. Left maxilliped I, lateral view. I. Left maxilliped II, lateral view. J. Left maxilliped III, lateral view. Scale = 1.6 mm (B, E, F, I), 2.2 mm (D, G, H, J), and 3.3 mm (A, C).

Fig. 93. Albunea marquisiana Boyko, 2000: A–F, Ƌ, 7.9 mm cl, USNM 268578, paratype; G, ♀, 5.2 mm cl, USNM 260948, paratype. A. Left pereopod I, lateral view. B. Left pereopod II, lateral view. C. Right pereopod III, lateral view. D. Right pereopod IV, lateral view. E. Abdominal somites I–VI, dorsal view. F. Telson of Ƌ, dorsal view. G. Telson of ♀, dorsal view. Scale = 1.1 mm (G), 1.6 mm (F), 3.3 mm (A–D), and 6.7 mm (E).

Fig. 94. Albunea holthuisi Boyko and Harvey, 1999: A, B, Ƌ, 8.1 mm cl, MNHN Hi­202, holotype; C, Ƌ, 7.7 mm cl, ANSP CA4644, paratype; D–J, Ƌ, 8.0 mm cl, MNHN Hi­204, paratype. A. Carapace, branchiostegite, and ocular peduncles, dorsal view. B. Ocular peduncles, dorsal view. C. Left antennule, lateral view. D. Left antenna, lateral view. E. Left mandible, mesial view. F. Left maxillule, lateral view. G. Left maxilla, lateral view. H. Left maxilliped I, lateral view. I. Left maxilliped II, lateral view. J. Left maxilliped III, lateral view. Scale = 1.1 mm (B), 1.6 mm (E, F, I), and 3.3 mm (A, C, D, G, H, J).

Fig. 95. Albunea holthuisi Boyko and Harvey, 1999: A, B, Ƌ, 8.0 mm cl, MNHN Hi­204, paratype; C, D, H, ♀, 8.1 mm cl, MNHN Hi­203, allotype; E–G, Ƌ, 8.1 mm cl, MNHN Hi­202, holotype. A. Left pereopod I, lateral view. B. Left pereopod II, lateral view. C. Left pereopod III, lateral view. D. Left pereopod IV, lateral view. E. Abdominal somites I–VI, dorsal view. F. Telson of Ƌ, dorsal view. G. Telson of Ƌ, lateral view. H. Telson of ♀, dorsal view. Scale = 1.7 mm (F, H) and 3.3 mm (A–E, G).