Leptostigma, Archibald & Rasnitsyn, 2023

Archibald, S. Bruce & Rasnitsyn, Alexandr P., 2023, Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands, Zootaxa 5278 (1), pp. 1-38 : 8

Leptostigma n. gen.

http://zoobank.org/ urn:lsid:zoobank.org:act:531F3657-F3D9-4A4D-86B7-1C146E74D627

Figs. 3A–D, 3G–K , 5–21 .

Type species. Leptostigma brevilatum n. sp.

Diagnosis. Distinguished from Cenocimbex by lacking dark triangular sclerite on R basal to pterostigma reaching 1-Rs+M near 1m-cu (absent or undetectable being small, pale); wing narrower (length/width 2.8 or more); cell 3rm long (2r-m to 3r-m distinctly longer than length of 3r-m); hind wing with cu-a well before middle of cell mcu; distinguished from Allenbycimbex by pterostigma narrow, maximum height about equal to height to C to R basal to pterostigma.

Species included. The type species and L. longiclava n. sp., L. longipallidum n. sp., L. longitenebricum n. sp., L. proxivena n. sp., L. fasciatum n. sp., and L. alaemacula n. sp.

Etymology. From the Greek leptos, meaning thin, and stigma, for the pterostigma. The name is neuter (see Examples below Art. 30.1.2, ICZN 1999).

Locality and age. Republic, Washington, United States of America, Tom Thumb Tuff Member of the Klondike Mountain Formation exposure A0307 ; and McAbee, British Columbia, Canada, Hoodoo Face beds; mid-Ypresian .

Figures

FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm.

FIGURE 5. Leptostigma brevilatum, holotype GSC 142839. A, photograph, and B–D, drawings: B, right forewing; C, left forewing; D, right hind wing. All to scale, 5 mm.

FIGURE 6. F-1182, tentative Leptostigma brevilatum, A, photograph, and B–E, drawings of: B, left forewing; C, right forewing; D, left hindwing, and; E, right hind wing. All to scale, 5 mm.

FIGURE 7. UWBM-54840, tentative Leptostigma brevilatum: A, photograph; B, drawing from part and counterpart. Scale bar 5 mm.

FIGURE 8. RBCM-EH2017-050-4628-001, tentative Leptostigma brevilatum, A, photograph, and B–C, drawings of the wings: B, right forewing; C, right hindwing.All to scale, 5 mm.

FIGURE 9. F-774, tentative Leptostigma brevilatum:A–B photographs and C–D, wing drawings:A, the whole fossil; B, closeup with arrow indicating the hind apical spur; C, right forewing; D, left forewing.A, C–D to 5 mm scale; B scale is 2 mm.

FIGURE 10. GSC 142840, tentative Leptostigma brevilatum: A, photograph, B, drawing of the forewing; scale is 5 mm.

FIGURE 11. Leptostigma longiclava holotype RBCM-EH2017.050.4085.001, A, photograph, B–E wing drawings: B, right forewing; C, left forewing; D, left hind wing; E, right hind wing. All to scale, 5 mm.

FIGURE 12. F-848, tentative Leptostigma longiclava, A, photograph, B–C, drawings: B, right forewing; C, left forewing. All to scale, 5 mm.

FIGURE 13. Leptostigma longipallidum, holotype GSC 142841, A, photograph, B–C, wing drawings: B, right forewing; C, left forewing. All to scale, 5 mm.

FIGURE 14. F-771, tentative Leptostigma longipallidum, A, photograph, B–E, wing drawings: B, right forewing; C, left forewing; D, right hind wing; E. left wing. All to scale, 5 mm.

FIGURE 15. F-1185 tentative Leptostigma longipallidum forewing: A, photograph; B, drawing. Scale is 5 mm.

FIGURE 16. Leptostigma longitenebricum holotype F-1565, A, photograph, B–C, wing drawings: B, left forewing; C, right forewing. All to scale, 5 mm.

FIGURE 17. F-1563, tentative Leptostigma longitenebricum, A, photograph and B–C, wing drawings: B, left forewing; C, right forewing. All to scale, 5 mm.

FIGURE 18. GSC 142842, tentative Leptostigma longitenebricum forewing: A, photograph; B, drawing. Scale bar 5 mm.

FIGURE 19. Leptostigma proxivena holotype F-1564, forewing: A, photograph; B, drawing. Scale bar 5 mm.

FIGURE 20. Leptostigma fasciatum holotype GSC 142843, forewing: A, photograph; B, drawing. Scale bar 5 mm.

FIGURE 21. Leptostigma alaemaculum holotype SR 92-18-08, forewing: A, photograph; B, drawing. Scale bar 5 mm.