Ptychoderes obsoletus Jekel, 1855

Mermudes, José Ricardo M. & Napp, Dilma Solange, 2006, Revision and cladistic analysis of the genus: Ptychoderes Schoenherr, 1823 (Coleoptera, Anthribidae, Anthribinae, Ptychoderini), Zootaxa 1182 (1), pp. 1-130 : 35-37

Ptychoderes obsoletus Jekel, 1855

Figs. 81 , 95, 109 , 123 , 169–172 , 215 , 256 , 265

Ptychoderes obsoletus Jekel, 1855: 61 ; Gemminger & Harold, 1872: 2722 (cat.); Bovie, 1906: 220 (cat.); Wolfrum, 1929: 6 (cat.); Rheinheimer, 2004: 10 (cat.).

Ptychoderes obsoleta [sic]; Blackwelder, 1947: 765 (cat.).

Male. Vestiture ( Fig. 215 ): pale dorsal vitta with dense, appressed, whitish scales intermingled with decumbent brownish to pale­brown scales denser on rostrum, and elytral interstriae 1, 2 and 4. Sides of pronotum, epipleura, apical declivity of elytra and pygidium with predominance of brown scales. Ventral body surface and legs with dense, appressed, yellowish­white scales, sparser on pro­ and mesosternum. Sides of prosternum, below lateral carina, with a continuous dark vitta of blackish­brown scales, slightly narrowed between pleural suture and posterior margin of prothorax. Sides of mesosternum with a narrow glabrous vitta. Mesepimera, metepisterna, sides of metasternum and ventrites with shallow setigerous punctures, each provided with a brownish scale. Outer surface of meso­ and metafemora with two spots of blackish­brown scales.

Rostrum 1.1–1.7 times as long as wide across base. Distance between scrobe and eyes about half or slightly more than maximum width of eye. Sides looking tricarinate because of careniform upper margin of scrobe; carinae somewhat oblique, upper one slightly thicker than lower carina, both weaker near margin of scrobe; with or without transverse carina between lower carina and margin of scrobal extension.. Scrobes careniform at dorsal and basal margins; extension wide, shallow with a deep distal pit. Mentum sparsely punctate­rugose; shallowly grooved at sides and at apical margin. Gula with dense, fine punctures. Frons with prominent carinae.

Antennae ( Fig. 215 ) either short, reaching elytral basal third, to moderately long, exceeding the middle of elytra. Segments IX and X weakly depressed dorsally; II one third shorter than III; IV to VI subequal in length; VII slightly longer; VIII–IX slightly shorter, subequal; X half as long as IX; XI one third longer than X (II = 0.6–0.9; III = 0.8–1.7; IV = 0.9–1.7; V = 0.8–1.8; VI = 0.7–1.8; VII = 0.7–2.1; VIII = 0.6–1.7; IX = 0.8–1.5; X = 0.5–0.7; XI = 0.8–1.2).

Prothorax ( Fig. 215 , 256 ) slightly longer than wide, feebly converging anteriorly from antebasal carina. Pronotal depression deep, divided by a prominent, more or less transverse tubercle reaching lateral margins of depression. Pronotum transversely rugose throughout, more so on tubercle and at sides near lateral carinae, less so inside depression behind tubercle. Antebasal carina widely interrupted at middle (at least twice width of carina), oblique towards lateral carina. Lateral carina strongly sinuous. Secondary carina straight, feebly interrupted at middle, not reaching lateral carina. Laterobasal carina reaching posterior margin of prothorax. Prosternum ( Fig. 256 ) raised, with coarse, deep, dense punctures; antecoxal furrow deep, moderately wide, curved in front of procoxae, partially interrupted at middle by a short prosternal projection almost as wide as the prosternal process at base. Mesosternum with moderately coarse, shallow, irregularly sparse punctures, and an irregularly transverse impression, deeper at middle near intercoxal process. Intercoxal process of mesosternum strongly impressed at apex, swollen at lateral and apical margins, without lateroapical projections. Elytra 2.2 times as long as wide across humeri, convex at apical third; interstriae 2 strongly depressed at basal two thirds; basal gibbosities weakly prominent; surface rugose from base to behind middle, more so at base, less so near suture. Sides slightly attenuate apically from basal third. Anteapical tubercles absent, interstriae 3, 5, 7 and 9 feebly swollen. Wings without radial cell.

Abdomen. Ventrite I with intercoxal process deeply grooved, sex patch absent. Ventrite V feebly emarginate at apex. Pygidium slightly longer than wide, depressed, with an elongate groove to fit elytra, almost rounded at apex.

Terminalia. Tergite VIII ( Fig. 81 ) slightly wider than long, convergent apically, rounded at apex; sternite VIII ( Fig. 95 ) strongly transverse, subparallel at sides, the apical margin somewhat truncate and acutely notched at middle, apodeme vestigial; apodeme of sternite IX ( Fig. 109 ) twice as long as arms. Tegmen ( Fig. 123 ) slightly emarginate at apex. Aedeagus ( Figs. 169–172 ) without arc between apodemes; tectum ( Fig. 170 ) straight at proximal margin, attenuate apically from middle; pedon ( Figs.169, 171 ) with body curved, weakly narrowed apically; apodemes curved. Internal sac ( Fig. 169 ) slightly shorter than aedeagus, spiculated medially, lateral membrane with pigmented rods; antemedian sclerites narrow and weakly sclerotized ( Fig. 172 ).

Female. Rostrum slightly shorter, 1.2 times as long as wide across base; extension of scrobe deep. Antennae shorter, exceeding the antebasal carina of prothorax; segments IX–X barely depressed dorsally at base; II to IV slightly increasing in length; V to VII subequal; VIII slightly shorter than VII; IX longer than VIII; X one third shorter than XI (II = 0.3–0.8; III = 0.4–0.7; IV = 0.5–0.9; V = 0.4–0.7; VI = 0.3–0.6; VII = 0.4–0.7; VIII = 0.3–0.6; IX = 0.5–0.9; X = 0.4–0.7; XI = 0.6–1.1). Elytra convex, slightly shorter than in male. Ventrite V as long as IV, truncate at apex. Terminalia similar to that of P. jordani , differing by the apical margin of both tergite VIII, obliquely truncate, and sternite VIII, truncate.

Measurements, in mm, male/female, respectively. TL = 12.7–23.5/11.0–20.9; RL = 1.7–3.4/1.6–2.7; RAW = 1.8–2.7/1.8–2.6; RBW = 1.5–2.0/1.3–2.2; HW = 1.2–1.8/ 1.0–1.6; DSO = 0.6–1.6/0.3–0.9; MLO = 1.2–1.8/1.0–1.6; PL = 3.5–5.7/3.2–5.4; PW = 3.5–5.2/3.1–5.3; EL = 8.5–13.7/7.2–13.6; EW = 3.7–6.1/3.3–6.3.

Type material

Not examined. Jekel (1855) did not mention how many specimens he examined, and according to Pitkin (2003) the type material is deposited in BMNH. Type locality: Brazil ( Jekel 1855).

Remarks

P. obsoletus is the sister group of P. magnus , by the median, well developed prosternal projection in front of the deep antecoxal furrow (25 1). They differ, besides the characters mentioned in the key, by the following features (condition present in P. magnus indicated in parenthesis): 1) antebasal carina of pronotum widely interrupted at middle, at least twice width of carina (narrowly interrupted at middle, less than twice width of carina); 2) secondary carina straight, weakly interrupted at middle, not reaching lateral carina (sinuous, reaching lateral carina near posterior margin of prothorax); 3) mesosternum with an irregular transverse impression, deeper at middle, near the intercoxal process (with deep, irregular, contiguous impressions); 4) intercoxal process of mesosternum strongly impressed at apex, lacking lateroapical projections (impressed at base, with lateroapical projections).

Distribution ( Fig. 265 ) BRAZIL (Pernambuco, Bahia and Espírito Santo).

Material examined

BRAZIL. Pernambuco: Pery­Pery (= Periperi ), 1 F, XII.1892, Gounelle leg. ( BMNH) . Bahia: 1 M, ex­coll. Castelnau­Roelofs. ( ISNB) . Espírito Santo: Conceição da Barra. 1 M, 1 F, XI.1968 , 4 M, 4 F, II.1969, Elias leg. ( DZUP, MZSP); Linhares ( Parq. Sooretama ), 1 M, V.1953, Teles leg. ( MNRJ), 1 M, 1 F, XI.1967, Oliveira leg. ( MNRJ) .

Figures

FIGURES 70–84. Eighth tergite, male, dorsal. 70, Hypselotropis annulicornis; 71, Ptychoderes mixtus; 72, P. bivittatus; 73, P. callosus; 74, P. rugicollis; 75, P. viridanus; 76, P. antiquus; 77, P. elongatus; 78, P.nebulosus; 79, P. longicollis; 80, P.jordani; 81, P. obsoletus; 82, P. magnus; 83, P. depressus; 84, P. virgatus; scale bar = 1mm.

FIGURES 85–112. 85–98, eighth sternite, male, ventral. 85, Ptychoderes mixtus; 86, P. bivittatus; 87, P. callosus; 88, P. rugicollis; 89, P. viridanus; 90, P. antiquus; 91, P. elongatus; 92, P. nebulosus; 93, P. longicollis; 94, P. jordani; 95, P. obsoletus; 96, P. magnus; 97, P. depressus; 98, P. virgatus. 99–112, ninth sternite, male, ventral. 99, Ptychoderes mixtus; 100, P. bivittatus; 101, P. callosus; 102, P. rugicollis; 103, P. viridanus; 104, P. antiquus; 105, P. elongatus; 106, P.nebulosus; 107, P. longicollis; 108, P. jordani; 109, P. obsoletus; 110, P. magnus; 111, P. depressus; 112, P. virgatus; scale bar = 1mm.

FIGURES 113–126. Tegmen, dorsal. 113, Ptychoderes mixtus; 114, P. bivittatus; 115, P. callosus; 116, P. rugicollis; 117, P. viridanus; 118, P. antiquus; 119, P. elongatus; 120, P. nebulosus; 121, P. longicollis; 122, P. jordani; 123, P. obsoletus; 124, P. magnus; 125, P. depressus; 126, P. virgatus. ap=apodeme; pf=preapical flange; sr=sclerotized ring; scale bar = 1mm.

FIGURES 158–176. Aedeagus. 158–161, Ptychoderes nebulosus, lateral, dorsal and ventral; 161, sclerite, dorsal; 162–164, P. longicollis, lateral, dorsal and ventral; 165–168, P. jordani, lateral, dorsal and ventral; 168, sclerite, dorsal; 169–172, P. obsoletus, lateral, dorsal and ventral; 172, sclerite, dorsal; 173–176, P. magnus, lateral, dorsal and ventral, 176, sclerite, dorsal; scale bar = 1mm.

FIGURES 215–220. 215, Ptychoderes obsoletus Jekel, 1855, male; 216, Ptychoderes magnus, Mermudes & Napp, 2004 male holotype; 217, Ptychoderes depressus Jekel, 1855, male; 218, Ptychoderes virgatus Fåhraeus, 1839, male; 219, Ptychoderes mixtus Jekel, 1855, male; 220, Ptychoderes bivittatus Jordan, 1894, male.

FIGURES 251–262. Prosternum. 251, Tribotropis prasinata; 252, Unanthribus grandis; 253, Ptychoderes virgatus; 254, P. jekeli, female holotype; 255, P. magnus, male holotype; 256, P. obsoletus; 257, P. longicollis, male holotype; 258, P. viridanus; 259, P. rugicollis; 260, P. callosus; 261, P.nebulosus; 262, P. elongatus.

FIGURES 265–266. 265. Geographical distribution of P. bivittatus (black squares), P. magnus (white triangles), P. obsoletus (black triangles). 266. Geographical distribution of P. mixtus (black squares), P. viridanus (lozenge), P. antiquus (white circles).