Chimarra vuda, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 28-30

Chimarra vuda , new species

Figs. 15 , 80–84

Chimarra vuda is similar to C. tokotaai and C. naitasirensis in having simple, dorsad-curved gonopods; a strongly enlarged phallotremal sclerite in the phallic apparatus; absence of large endothecal spines; and an endotheca with a group of minute, strongly sclerotized spicules. Chimarra vuda is distinguished from the other 2 species by its nearly parallel-sided gonopods in lateral view; a phallotremal sclerite that is about 2x longer than the width of the posterior part of phallotheca; and by the presence of 3 groups of sclerotized spicules in the phallic endotheca.

Male. Body yellowish-brown, dorsal part of meso- and metathorax brown. Large dark area between lateral and anterior ocelli. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 15 ). Forewings 6.6 mm (n=1), brown; broad, ratio of length to breadth 3.3; R1 slightly undulating before crossvein r; radial sector slightly produced anterad immediately before discoidal cell; discoidal cell originating slightly distally of mid-length of wing, nearly 3x longer than wide; median cell slightly shorter than discoidal cell; crossvein r originating from basis of fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork III 1 /5th as long as wing; fork V about as long as fork I, shorter than fork II; Cu2 situated close to A at wing margin. Hind wings 5.3 mm (n=1), brown; broad, ratio of length to breadth 2.9; margin slightly incurved at arculus, where Cu1 and Cu2 fused with margin; fork III 2x longer than discoidal cell and 1/6th as long as wing; fork V about as long as fork I; 1A+2A 4x longer than 1A.

Male genitalia ( Figs. 80–84 ). Segment IX about as long as high; each anterodorsal margin nearly straight in lateral view; each ventral margin nearly straight; each posterior margin slightly produced posterad into rounded, triangular lobe starting at mid-height of cercus; ventral side of posterior-most part of segment IX with scattered setae ( Fig. 82 ). In dorsal view with narrow anterior lobes; anterodorsal margin with wide, deep incision in dorsal view. In ventral view segment IX with shallowly concave anterior margin and almost straight posterior margin; posterior margin without central projection. Tergum X simple; tapering apically in lateral view ( Fig. 80 ); orienting ventrad, nearly straight; divided into 2 slightly mesad-curved, lateral branches from base in dorsal and ventral view ( Figs. 81, 82 ); each branch uniformly narrowing posteriorly in dorsal and ventral view, apically pointed, without processes; sensillae invisible. Cerci large, broad, located immediately below dorsal margin of segment IX and tergum X and oriented posterodorsad in lateral view; tube-shaped in dorsal view; covered by long setae. Gonopodsunbranched, longer than segment IX, slightly curved dorsad and sickle-shaped in lateral view; anterodorsal margin shallowly concave; ventral margin uniformly convex; posteroventral corner not produced into tooth in lateral view; in ventral view forming broad plates widening posteriorly to 1/2-length; tapering from 1/2-length into mesad curved apex; each basal 1/2 of mesal margin nearly straight, weakly diverging with minute mesal tooth. Phallic apparatus nearly 2x longer than rest of genitalia: phallotheca, in lateral and ventral view, with anterior part slightly more than 2x thicker than posterior part; apicoventral spine absent on phallotheca; strongly enlarged phallotremal sclerite in phallic apparatus forming complicated structure 1/3rd as long as phallic apparatus; large endothecal spines absent; endotheca with 3 groups of minute, strongly sclerotized, well separated spicules; anterior group very densely packed, dark, located ventrally of posterior part of phallotremal sclerite; apical 2 groups formed by densely packed spicules, and more scattered spicules.

Female. Unknown.

Holotype male: VITI LEVU: Vuda Prov., Koroyanitu N.M.P. Abaca Village, Malaise trap, 6–26.v.2003, 17°40’S, 177°33’E [17.3333°S, 177.55°E], 400 m, leg. M. Irwin, E. Schlinger & M. Tokota'a [loc#01] [ FNIC].

Etymology: Vuda , named after the Vuda Province, the type locality of the species.

Distribution: Viti Levu.

Figures

FIGURES 14 – 21. Right wings, dorsal views. 14 — C. tokotaai, new species, holotype; 15 — C. vuda, new species, holotype; 16 — C. naitasirensis, new species, holotype; 17 — C. veisarensis, new species, holotype; 18 — C. fijiana, new species, holotype; 19 — C. abacensis, new species, holotype; 20 — C. manni Banks, non-type; 21 — C. cakaudrovensis, new species, holotype.

FIGURES 80 – 84. Chimarra vuda, new species, holotype. 80 — genitalia, left lateral view; 81 — genitalia, dorsal view; 82 — genitalia, ventral view; 83 — phallic apparatus, left lateral view; 84 — phallic apparatus, ventral view.