Chimarra malickyi, Johanson, Kjell Arne & Oláh, János, 2012

Johanson, Kjell Arne & Oláh, János, 2012, Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species, Zootaxa 3354, pp. 1-58 : 53-55

Chimarra malickyi , new species

Figs. 31 , 165–169

Chimarra malickyi is most similar to C. psychodida , particularly in the shape of the gonopods in lateral view. It is distinguished from C. psychodida by the narrower anteroventral lobe of segment IX in lateral view, longer tergum X, and absence of megasetae on the apex of the dorsal branch of each gonopod.

Male. Body yellowish brown, dorsal part of meso- and metathorax brown. Large dark area between lateral and anterior ocelli. Foreleg anterior claw as long as foreleg spur.

Wings ( Fig. 31 ). Forewings 4.9 mm (n=1), brown; broad, ratio of length to breadth 3.4; R1 curved before crossvein r; radial sector weakly produced anterad immediately before discoidal cell; discoidal cell originating at mid-length of wing, about 3x longer than wide; median cell slightly longer than discoidal cell; crossvein r situated immediately basally of fork I; fork I originating before crossvein s at distance equal to length of crossvein s; nygma located near base of fork II; fork III 1 /6th as long as wing; fork V as long as fork I, shorter than fork II; Cu2 nearly fused with A at wing margin. Hind wings 4.0 mm (n=1), brownish-grey; broad, ratio of length to breadth 3.0; margin weakly incurved at arculus, where Cu1 and Cu2 fused with margin; fork III nearly 1.5x longer than discoidal cell and 1/7th as long as wing; fork V as long as fork I; 1A+2A 4x longer than 1A.

Male genitalia ( Figs. 165–169 ). Segment IX slightly shorter than high; ventral part of segment produced into hyperbolic lateral plates in lateral view, with minute ventral projection at anteroventral corner; dorsal part of segment produced into small rounded, anterad directed lobe, finger-like in dorsal view; each anterodorsal margin deeply concave in lateral view; each ventral margin convex; each posterior margin forming slightly convex plate originating immediately below cercus; posterior quarter of ventrum IX with setae ( Fig. 167 ). In dorsal view anterodorsal margin forming wide incision with short, narrow, anterad-orienting processes on each side. In ventral view segment IX with nearly parallel lateral margins, with deeply concave anterior margin and nearly straight posterior margin; posterior margin without central projection. Tergum X simple; forming pair of lateral branches from basis of segment; lateral branches slightly curved ventrad in lateral view, with small tooth on dorsal margin at 2/3rds its length; in dorsal view straight, diverging along their length, uniformly narrow before tapering into finger-like apex. Cerci short, cylindrical, situated on posterior margin of segment IX at mid-height of basis of tergum X in lateral view and oriented posterodorsad in lateral view; covered by long setae. Gonopods nearly as long as segment IX, 3- branched. Dorsal branch of each gonopod long, but not exceeding tergum X in lateral view, uniformly slender, straight before slightly ventrad curved apex in lateral view; anterodorsal margin nearly straight before apex, ventral margin straight before concave at apex; in dorsal view slightly converging long their length; without megasetae. Ventral branch of each gonopod strongly produced ventrad in lateral view, with undulating margins; in right angle to dorsal brach; in ventral view, ventral branches forming broad plates slightly curved mesad, with megasetae along posterior margin. Mesal branches simple, darkly pigmented, about 2x longer than cercus; in lateral view orienting posterodorsad; in ventral view thorn-shaped, apices pointing mesad, nearly tangential. Phallic apparatus about 1.5x longer than rest of genitalia; curved along its length: phallotheca, in lateral view, with anterior part about 4x thicker than posterior part; and in ventral view anterior part about 3x wider than posterior part; apicoventral spine absent on phallotheca; large phallotremal sclerite in phallic apparatus forming U-like structure with central spine, about as long as endothecal spines; 2 short, nearly black endothecal spines present.

Female. Unknown.

Holotype male: VITI LEVU: Naitasiri Prov., 3.3 km N Veisari, logging rd. to Waivudava, Malaise trap, 8–31.iii.2003, 18.0592°S, 178.367°E, 300 m, leg. M. Tokota’a [loc#20] [ FNIC].

Paratype: Vuda Prov., Koroyanitu Pk., 1 km E Abaca Vlg., Savuione Trail, Malaise trap, 26.xi–3.xii.2002, 17.667°S, 177.55°E, 800 m, leg. M. Irwin, E. Schlinger & M. Tokota’a [loc#03] — 1 male [ NHRS, DNA voucher IM9].

Etymology: Malickyi , named after Hans Malicky, in recognition of his great contribution to our knowledge of caddisflies.

Distribution: Taveuni and Vanua Levu.

Figures

FIGURES 30 – 31. Right wings, dorsal views. 30 — C. levuensis, new species, holotype; 31 — C. malickyi, new species, holotype.

FIGURES 165 – 169. Chimarra malickyi, new species, holotype. 165 — genitalia, left lateral view; 166 — genitalia, dorsal view; 167 — genitalia, ventral view; 168 — phallic apparatus, left lateral view; 169 — phallic apparatus, ventral view.