Thyropygus carli Attems, 1938

Pimvichai, Piyatida, Enghoff, Henrik & Panha, Somsak, 2011, A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae), Zootaxa 2980, pp. 37-48 : 39-41

Thyropygus carli Attems, 1938

( Figs. 1 A–D)

Thyropygus cuisinieri carli Attems, 1938: 281 ; Hoffman 1975: 127 Thyropisthus cuisinieri carli: Attems 1942: 79 ; Demange 1961: 124 Thyropisthus (Duplopisthus) cuisinieri carli: Demange 1983: 563 Thyropygus carli: Enghoff 2004: 36

Material examined. SYNTYPES (2 vials) VIETNAM (S-Annam), Nhatrang, 12° 14ˏ 43˝ N, 109° 11ˏ 57˝ E. C. Dawydoff leg., ( NMW 2558, NMW 2559). 2 males, 4 females VIETNAM, Nhatrang, 12° 14ˏ 43˝ N, 109° 11ˏ 57˝ E. 22 March 2000. P. Gravlund and A. R. Rasmussen leg., ( ZMUC). 3 males, 1 female VIETNAM, Nhatrang, Bao Dai’s village hotel, 12° 12ˏ 39˝ N, 109° 12ˏ 59˝ E. 9–13 March 2004. J. B. Rasmussen leg., ( ZMUC). 2 males VIETNAM, Cauda, near Nhatrang, 12° 15ˏ 4˝ N, 109° 11ˏ 45˝ E. October, 27 November 1959. J. Knudsen leg., ( ZMUC). 1 male, 1 juvenile S. VIETNAM, Phantiet, 12° 14ˏ 31˝ N, 109° 11ˏ 14˝ E. 2002. P. Gravlund leg., ( ZMUC). 1 male bought in pet shop ( ZMUC).

Diagnosis. A species of the cuisinieri subgroup. Lateral process of posterior coxal fold (plp) huge, thumblike. Similar in this respect to T. cuisinieri and T. foliaceus . Differs from all other species of the T. cuisinieri subgroup by having two short, straight, broadly triangular distal processes on the anterior coxal fold (ac).

Description. Adult males with 54–61 podous rings, no apodous rings. Length 11–12 cm, width 8.0– 8.9 mm. Adult females with 57–60 podous rings, no apodous rings. Length 9–12 cm, width 7.3–9.4 mm. Color in life unknown; preserved specimens with antennae, legs pale brown; metazona dark brown; prozona whitish brown; epiproct, paraprocts and hypoproct brown.

Gonopods ( Figs. 1 A–D): Anterior coxal fold (ac) ( Fig. 1 A) basally slender, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two short, straight, broadly triangular distal processes, the lateral one produced into a short, sharp spine, pointing distad, the mesal one broad, apically pointed, pointing distad. Posterior coxal fold (pc) ( Fig. 1 B) basally with relatively high lateral paracoxites (px); mesal process (pmp) very small, short, rounded lamellar, directed distolaterad; lateral process (plp) huge, thumblike, directed distolaterad, covering pmp in posterior view. At the base of plp, a blunt ridge (br). Telopodite ( Figs. 1 C–D) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) long, curving mesad in the horizontal plane, distinctly crenulated along inner curvature, in situ resting against plp, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a slender, sharply pointed, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about six to eight brownish blepharochaetae (bp).

Figures

FIGURE 1. Thyropygus carli (specimen from Nhatrang), gonopods. A: anterior view, left telopodite removed. B: posterior view, left telopodite removed. C: left telopodite, posterior-mesal view. D: left telopodite, anterior-lateral view.