Daphniola magdalenae Falniowski

Falniowski, Andrzej & Sarbu, Serban, 2015, Two new Truncatelloidea species from Melissotrypa Cave in Greece (Caenogastropoda), ZooKeys 530, pp. 1-14 : 5

Taxon classification Animalia Littorinimorpha Hydrobiidae

Daphniola magdalenae Falniowski sp. n.

Types.

Ethanol-fixed specimens, Melissotrypa Cave, Thessalia, Greece, 39°52'38"N, 22°02'58"E, sulphidic lake, near the shore, June 2014, S. Sarbu coll., holotype: ZMUJ-M.2109; 20 paratypes: ZMUJ-M.2110-ZMUJ-M.2130.

Diagnosis.

Shell relatively big, valvatiform-trochiform; soft parts with no pigment, no eyes, penis with long and slender filament and big outgrowth on the left side. Readily distinguished from geographically and closely related Daphniola exigua (= Daphniola graeca ) by its bigger size (2.5 vs. 1.5 mm), reddish operculum, broader base and longer and thinner filament of the penis. Differentiated from Daphniola louisi (from Kessariani at Athens) by its larger size, higher spire, longer and thinner filament and more prominent outgrowth on the left side of the penis. Differs from Daphniola hadei (from Gythion at Peloponnese) by its double size, higher spire and much more prominent outgrowth on the left side of the penis.

Description.

Shell (Fig. 2 A–D) valvatiform-trochiform, up to 2.68 mm tall, having 3.5-3.75 whorls, spire height 16% height of shell, and 13-16% width of shell. Teleoconch whorls moderately convex, evenly rounded, growing rapidly in diameter. Aperture circular, parietal lip complete, umbilicus very broad, outer lip simple, orthocline. Teleoconch with delicate growth lines, periostracum pinkish. Shell parameters for a series of paratypes are given in Table 2. On the surface there are numerous pellets of sediment, most probably of sulfuric bacteria.

Inner and outer sides of operculum smooth. Operculum pink (Fig. 2 A–D). Protoconch of 1.25-1.40 whorls growing slowly (Fig. 3), with a net-like pattern of dense depressions, their shape irregular (Fig. 4), covering all the protoconch and initial part of the teleoconch.

Radula (Figs 5-7): taenioglossate, typically hydrobiid; the cusps on the central, lateral and inner marginal teeth prominent, long and sharp; the central tooth trapezoid (Figs 5-6), with one pair of big basal cusps arising from the tooth face (Fig. 5) and numerous long cusps along the cutting edge, the basal tongue broadly V-shaped and about equal in length to the lateral margins, lateral cusps five–six. Lateral teeth (Figs 6-7) having four cusps on inner, and five cusps on outer side, central cusp broad and blunt. Inner marginal tooth (Fig. 7) with 35-36 cusps, outer marginal teeth (Figs 6-7) with 21-23 cusps.

Animal brownish, with no pigment, and no eyes (Fig. 8). Penis (Figs 9-11) having broad base bent U-shaped in natural position (Fig. 8), long and narrow filament and prominent outgrowth on its left edge. Female reproductive organs (Fig. 12) with big bursa copulatrix with long duct and two small receptacula seminis.

Etymology.

Named in memory of Dr Magdalena Szarowska, a malacologist, wife and best friend of the first author.

Distribution and habitat.

Known from the type locality only.