Baradina

Chandler, Donald S., 2023, The Eupsenius LeConte of North America and the Lesser Antilles with Notes on the Brachyglutini Subtribes Eupseniina Park and Baradina Park (Coleoptera: Staphylinidae: Pselaphinae), The Coleopterists Bulletin 77 (3), pp. 397-412 : 399-400

Subtribe Baradina

Specimens Examined. Euphalepsus , 25+ species, cleared specimens of two species; Phalespoides , five species at hand, cleared specimens of two species; Eupsenina , seven species at hand, cleared specimens of two species. Eupsenina Raffray is here transferred to the subtribe Baradina .

Diagnosis. Body with hindbody (meso-, metatho- rax + abdomen) swollen, often almost globular; forebody (head + prothorax) narrow when compared to hindbody. Head often noticeably elongate, extending well beyond antennal insertions, with long tempora, clypeofrontal area depressed to apex, lacking distinct rostrum, antennal tubercles prominent, rounded, positioned dorsal to anterior margin of eyes or dorsal to eyes; often with distinct apicolateral genal projections, may be reduced to small lobe, elongate thin setose lobe, or spinose projection; lacking lateral ocular-maxillary carinae, gula with median longitudinal impression with distinct to faint median carina/ridge; mandibles with single lateral seta ( Euphalepsus , Phalespoides ) or lacking lateral seta ( Eupsenina ); antenna geniculate; maxillary palpi typical in form but comparatively small, with four palpomeres. Mesoventral area: lateral mesoventral foveae small, widely separated, lacking median mesoventral fovea. Elytra lacking lateroapical cleft, area broadly rounded. Metacoxae contiguous/ subcontiguous for Euphalepsus and Phalespoides , widely separate for Eupsenina , metacoxae flat except for short projection for trochanteral articulation, projection not prominent. Anterior tarsal claw on each leg small in Euphalepsus , very small and variably present for Phalespoides , absent for Eupsenina . Tergites lacking free paratergites, tergite 1 with inner and outer carinae at position of paratergite margins, tergites 1–3 and ventrites 2–4 subequal in length, abdomen approximately cylin- drical.Aedeagal form more similar to that of genera of Panaphantina, subtribe of Euplectitae.

Sergei Kurbatov and Giorgio Sabella are working on this group, and commented earlier ( Kurbatov and Sabella 2015) that the Baradina does not belong in the Brachyglutini, but did not place the subtribe elsewhere as further study was required for a decision based on a thorough analysis, a decision with which I certainly concur. Eupsenina is the odd member of the Baradina with widely separated metacoxae (analagous to Eupsenius ), prominent apicolateral genal projections, geniculate antennae with the apical two antennomeres moderately setose, and antennomere XI long and thin, apex of XI sometimes penicillate ( Fig. 13 , undescribed Eupsenina species). The dorsum has sparse, short, appressed setae, and is further differentiated from Eupsenius on that basis and by the major differences of the head form (extended head lacking rostrum and the gular modifications).

Figures

Figs. 6–14. Eupsenius species, dorsal habitus photos and E. politus distribution map. 6) E. dilatatus; 7) E. glaber; 8) E. peckorum, new species; 9) E. rileyi, new species; 10) E. politus. Eupsenina raffrayi: 11) Dorsal habitus; 12) Anterior view. 13) Eupsenina undescribed species, dorsal habitus. 14) Distribution of Eupsenius politus.