Scybalocanthon maculatus ( Schmidt, 1920 ),

Silva, Fernando A. B. & Valois, Marcely, 2019, A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini), Zootaxa 4629 (3), pp. 301-341: 324-326

publication ID

https://doi.org/10.11646/zootaxa.4629.3.1

publication LSID

lsid:zoobank.org:pub:1D47B452-0A40-4BC5-9BBA-C06DEB298C9A

persistent identifier

http://treatment.plazi.org/id/9D12E557-3A4E-FF87-95F1-FF32FD2EC62D

treatment provided by

Felipe

scientific name

Scybalocanthon maculatus ( Schmidt, 1920 )
status

 

Scybalocanthon maculatus ( Schmidt, 1920) 

( Figs. 1View FIGURE 1 N–O, 4N, 5M, 6B)

Canthon maculatus Schmidt, 1920: 127  (original description); Schmidt 1922: 65, 77 (diagnosis, distribution); Balthasar 1939: 193 (identification key); Krajcik 2006: 28 (catalog); Krajcik 2012: 64 (catalog); Vaz-de-Mello & Cupello 2018: 45 (data of type specimens), 46 (figs. 29–30).

Canthon maculatum: Blackwelder 1944: 200  (checklist).

Scybalocanthon maculatus: Pereira & Martínez 1956: 115  , 119 (catalog and identification key); Vulcano & Pereira 1964: 638 (catalog); Vulcano & Pereira 1967: 555 (distribution, identification key); Halffter & Martínez 1977: 68 (checklist); Carvajal et al. 2011: 316 (cited for Ecuador); Chamorro et al. 2018: 98 (cited for Ecuador); Chamorro et al. 2019: 230 View Cited Treatment , 231 (catalog of dung beetles species from Ecuador).

Scybalocanthon maculatum  : Vaz-de-Mello 2000: 195 (checklist).

Diagnosis. Specimens of S. maculatus  are readily distinguishable from all other species by the pronotum usually with three spots along midline (anterior spot triangular in shape; posterior spots rounded in shape) and one rounded spot on each side of midline ( Fig. 1NView FIGURE 1). The three central spots can be fused ( Fig. 1OView FIGURE 1). In some individuals from Loja ( Ecuador), spots are missing. In addition, males have a diagnostic aedeagus with parameres elongate, not excavated ventrally ( Fig. 4NView FIGURE 4); and endophallus lacking bristles or microbristles close to the FLP sclerite ( Fig. 5MView FIGURE 5).

Description. Body. Oval, lateral edges rounded. Surface opaque, completely microgranulate, with small and dense punctures difficult to see under low magnification. Color. Most of pronotum, lateral portion of hypomera, lateral portion of abdominal ventrites, pygidium, and middle of femora yellow. Head, spots of pronotum, elytra, internal portion of hypomera, prosternum, mesoventrite, mesoepisternae, metaepisternae, metaventrite, central portion of abdominal ventrites, proximal and distal portions of femora, and tibiae black. Length. 7.8–8.7 mm. Thorax. Anterior angles of pronotum acute (approximately 80°). Lateral margin regularly curved outward, not forming an angle at the middle portion. Elytra. Striae thin and shiny, punctures inconspicuous. Eighth stria with a thin carina at the anterior portion that disappears after the carina and reappears at the middle of elytra. Aedeagus. Parameres slightly asymmetrical, with subrectangular shape, truncate apically ( Fig. 4NView FIGURE 4). Dorsal margin of parameres slightly curved inward from the basal to medial portions and substraight at the apical portion. Ventral margin of parameres substraight. SRP circular, with substraight handle-shaped extension ( Fig. 5MView FIGURE 5). FLP I-shaped ( Fig. 5MView FIGURE 5). A+SA with two superposed and elongate sclerites ( Fig. 5MView FIGURE 5). AS I-shaped ( Fig. 5MView FIGURE 5).

Type material. Canthon maculatus Schmidt  , lectotype ♂ (here designated; NHRS)  “ Santa Jnéz / ( Ecuad. ) / R. Haensch S.” // “ Typus ” // “ Typ. ” // “9846 / E92 +” // “maculatus / Type m.” // “ LECTOTYPE ♂ / Canthon  / maculatus  / Schm. / des. F. Z. Vaz-de-Mello ”  . Paralectotypes 2♂ ( NHRS): “Santa Jnéz / (Ecuad.) / R. Haensch S.” // “9843 / E92 + or 9842 / E92 +” // “ Scybalocanthon  / maculatus  / (Schm) / P. PEREIRA DET.” // “PARACTO- TYPE / Canthon  / maculatus  / Schm. / des. F. Z. Vaz-de-Mello ”  . Paralectotype ♀ ( NHRS)  : “Santa Jnéz/ (Ecuad.)/ R. Haensch S.” // “9845 / E92 +” // “PARACTOTYPE / Canthon  ♀ / maculatus  / Schm. / des. F. Z. Vaz-de-Mello ”. Paralectotype  ♀ ( NHRS)  : “ Santa Jnéz / (Ecuad.) / R. Haensch S.” // “29845 / E92 +” // “26 / 56 ” // “PARACTO- TYPE / Canthon  / maculatus  / Schm. / des. F. Z. Vaz-de-Mello ”. Paralectotype  3♂ and 1♀ ( SMTD)  : “ Ecuador / Baron” // “ Coll. C. Felsche / Kauf 20 / 1918” // “PARACTOTYPE / Canthon  ♂ / maculatus  / Schm. / des. F. Z. Vaz-de-Mello ”. Paralectotype  ♂ ( SMTD)  : “ Ecuador / 5255” // “ Canthon  / maculatus  / a. Schmidt ” // “PARAC- TOTYPE / Canthon  ♂ / maculatus  / Schm. / des. F. Z. Vaz-de-Mello ”. Paralectotype  ♀ ( SMTD)  : “ Santa Jnéz / ( Ecuad. ) / R. Haensch S.” // “ Canthon  / maculatus  / a. Schmidt ” // “ Coll. C. Felsche / Kauf 20/ 1918” // “ Typus ” // “PARACTOTYPE / Canthon  ♀ / maculatus  / Schm. / des. F. Z. Vaz-de-Mello ”. Paralectotype  1♂ and 1♀ ( SMTD)  : “ Santa Jnéz / (Ecuad.)/ R. Haensch S.” // “ Coll. C. Felsche / Kauf 20/ 1918”. // “PARACTOTYPE / Canthon  ♀ or ♂ / maculatus  / Schm. / des. F. Z. Vaz-de-Mello ”  .

Non-type material [90 males, 92 females]: ECUADOR: NAPO: Río Hollin (77°40’W, 00°42’S), 6.xii.1997, 1100 m, Z. Aguilar— 1♂ ( CEMT); Río Hollin, C. Padilha— 1♂ ( CEMT); Río Hollin (77°45’W, 00°57’S), J. C. Santos— 1♂ 1♀ ( CEMT); El Reventador, 6.viii. 1998, 760 m, C. Caplo— 1♀ ( CEMT); Los Guacanayos, Piviyacu, 29–31.xii.1995, P. Padilha— 1♀ ( CEMT); La Merced de Jondachi, Rio Jondachi, 25.viii.2010, W. Chamoro— 1♂ ( CEMT); km 11.1 Sarayacu-Loreto road, 1200 m (0°42’37’’S, 77°43’10’’W), 14.vii.1994, F. Génier, cloud forest, carrion trap— 9♂ 7♀ ( CMNC); km 11.1 Sarayacu-Loreto road, 1200 m (0°42’37’’S, 77°43’10’’W), 20.vii.1994, F. Génier, cloud forest, feces trap— 9♂ 5♀ ( CMNC); km 25.4 Sarayacu-Loreto road, 950 m (0°43’6’’S, 77°37’18’’W), 12.vii.1994, F. Génier, cloud forest, feces trap— 4♂ 3♀ ( CMNC); km 25.4 Sarayacu-Loreto road, 950 m (0°43’6’’S, 77°37’18’’W), 14.vii.1994, F. Génier, cloud forest, feces trap— 4♂ 6♀ ( CMNC); km 7.3 Sarayacu-Loreto road, 1200 m (0°43’2’’S, 77°44’11’’W), 11.vii.1994, F. Génier, cloud forest, feces trap— 3♂ 1♀ ( CMNC); km 7.3 Sarayacu- Loreto road, 1200 m (0°43’2’’S, 77°44’11’’W), 14.vii.1994, F. Génier, cloud forest, feces trap— 4♂ 3♀ ( CMNC); km 7.3 Sarayacu-Loreto road, 1200 m (0°43’2’’S, 77°44’11’’W), 20.vii.1994, F. Génier, cloud forest, feces trap— 1♂ 2♀ ( CMNC); PASTAZA: Mera, Estação Biológica Pinto Mirador UTE [Usina Termeletrica] (78°06’29”W, 1°27’25”S), 22.iii.2012, 1000 m, C. Luzuriaga— 1♂ 1♀ ( CEMT); 1 km E Mera, 1100 m (1°27’12’’S, 78°6’31’’W), 13–17.vii.1976, S. Peck, ravine edge forest, carrion & dung traps— 1♂ ( CMNC); Llandia, 17 km N Puyo, 1000 m (1°21’3’’S, 77°58’4’’W), 16.vii.1994, F. Génier, remnant rainforest, hand collecting— 1♀ ( CMNC); Llandia, 17 km N Puyo, 1000 m (1°21’3’’S, 77°58’4’’W), 19.vii.1994, F. Génier, remnant rainforest, feces trap— 2♂ 1♀ ( CMNC); TUNGURAHUA: Baños, El Topo (78°22’52”W, 1°23’41”N), 23.i.2011, 1590 m, G. Maldonato— 15♂ 16♀ ( CEMT); 4.3 km E Río Negro, remnant cloud forest, 18.vii.1994, 1200 m, F. Génier— 2♂ 1♀ ( CEMT). 3 km W Río Negro, 1200 m (1°24’36’’S, 78°14’29’’W), 18.vii.1994, F. Génier, cloud forest, feces trap— 2♀ ( CMNC); 4.3 km E Río Negro, 1200 m (1°26’8’’S, 78°10’37’’W), 18.vii.1994, F. Génier, remnant cloud forest, feces trap— 8♂ 18♀ ( CMNC); 4.3 km E Río Negro, 1200 m (1°26’8’’S, 78°10’37’’W), 20.vii.1994, F. Génier, cloud forest, feces trap— 13♂ 13♀ ( CMNC, MZUFPA); 6 km E Río Negro, 1500 m (1°26’53’’S, 78°10’13’’W), 17.vii.1973 – 13.vii.1976, S. Peck, carrion trap (traps 49-50)— 6♂ 5♀ ( CMNC); 8 km E Río Negro, 10 km W Pastaza [= Shell City], 1400 m (1°26’40’’S, 78°10’28’’W), 13–17.vii.1976, S. Peck, forest, carrion trap— 1♀ ( CMNC); 8 km E Río Negro, 10 km W Pastaza [= Shell City], 1400 m (1°26’40’’S, 78°10’28’’W), 13–17.vii.1976, S. Peck, forest, dung traps— 3♂ ( CMNC); LOJA, Loja (3°59’30’’S, 79°11’36’’W), [no date], Abbé Gaujon— 2♀ ( CMNC); MORONA- SANTIAGO, Ángel Rouby, sitio 8, 1300 m (2°21’39’’S, 78°2’2’’W), 2.ii.2002, J. Celi, pitfall trap, feces— 1♀ ( CMNC).

Distribution. Known from Ecuador ( Fig. 6BView FIGURE 6). Endemism areas: Brazilian sub-region: Boreal Brazilian dominion: Napo province (see Morrone 2014; fig. 12).

NHRS

Swedish Museum of Natural History, Entomology Collections

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Scybalocanthon

Loc

Scybalocanthon maculatus ( Schmidt, 1920 )

Silva, Fernando A. B. & Valois, Marcely 2019
2019
Loc

Canthon maculatus

Vaz-de-Mello, F. Z. & Cupello, M. 2018: 45
Krajcik, M. 2012: 64
Krajcik, M. 2006: 28
Balthasar, V. 1939: 193
Schmidt, A. 1922: 65
Schmidt, A. 1920: 127
Loc

Canthon maculatum:

Blackwelder, R. 1944: 200
Loc

Scybalocanthon maculatus: Pereira & Martínez 1956: 115

Chamorro, W. & Marin-Armijos, D. & Asenjo, A. & Vaz-de-Mello, F. Z. 2019: 230
Chamorro, W. & Marin-Armijos, D. & Granda, V. & Vaz-de-Mello, F. Z. 2018: 98
Carvajal, V. & Villamarin, S. & Ortega, A. M. 2011: 316
Halffter, G. & Martinez, A. 1977: 68
Vulcano, M. A. & Pereira, F. S. 1967: 555
Vulcano, M. A. & Pereira, F. S. 1964: 638
Pereira, F. S. & Martinez, A. 1956: 115
Loc

Scybalocanthon maculatum

Vaz-de-Mello, F. Z. 2000: 195