Scybalocanthon papaxibe Silva & Valois, 2019

Silva, Fernando A. B. & Valois, Marcely, 2019, A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini), Zootaxa 4629 (3), pp. 301-341 : 330

publication ID	https://doi.org/ 10.11646/zootaxa.4629.3.1
publication LSID	lsid:zoobank.org:pub:1D47B452-0A40-4BC5-9BBA-C06DEB298C9A
DOI	https://doi.org/10.5281/zenodo.4335184
persistent identifier	https://treatment.plazi.org/id/9D12E557-3A44-FF83-95F1-F8D9FB7DC361
treatment provided by	Felipe
scientific name	Scybalocanthon papaxibe Silva & Valois
status	sp. nov.

Treatment

Scybalocanthon papaxibe Silva & Valois View in CoL , new species

( Figs. 1T View FIGURE 1 , 4R View FIGURE 4 , 5Q View FIGURE 5 , 6D View FIGURE 6 )

Etymology. The new specific name, “papaxibe” [papa = eater; xibé = cassava flour porridge], is a typical term in the Pará state in Brazil. This term is often attributed to people who are born in Pará due to the old habit of consuming a type of cassava flour porridge. The specific name should be treated as a Latin adjective.

Diagnosis. Specimens of S. papaxibe are similar to those of S. arnaudi , S. federicoescobari , S. martinezi , S. pinopterus , and S. uniplagiatus by having the pronotum uniformly colored, or with one elliptical spot on the central portion ( Figs. 1T View FIGURE 1 , 2A View FIGURE 2 , 1P View FIGURE 1 , 2K View FIGURE 2 , 1D, 1I View FIGURE 1 ); femora almost completely yellow or brown, with black spots only on the tips ( Fig. 3G View FIGURE 3 ); eighth elytral stria with thin carina on anterior portion ( Fig. 3D View FIGURE 3 ); endophallus with bristles right beside the FLP sclerite ( Figs. 5Q, N, W, D, H View FIGURE 5 ); and additional sclerite ( AS) absent ( Figs. 5Q, N, W, D, H View FIGURE 5 ). They can be distinguished from those of S. pinopterus and S. arnaudi by the strongly asymmetrical parameres; left paramere with acute projection on dorsal margin and bilobate excavation on ventral margin ( Fig. 4R View FIGURE 4 ) (other species with slightly asymmetrical parameres, lacking acute projection and bilobate excavation on dorsal and ventral margins, respectively ( Figs. 4S, E View FIGURE 4 )); from those of S. uniplagiatus , S. martinezi , and S. federicoescobari by the bilobate excavation on the ventral margin of the left paramere wider than deep in lateral view, extending along one-fourth of the paramere margin in lateral view ( Fig. 4R View FIGURE 4 ) (in S. martinezi the bilobate excavation is wider and deeper, extending at least one-half of the paramere margin ( Fig. 4O View FIGURE 4 ); in S. uniplagiatus its excavation extending along one-third of the paramere margin ( Fig. 4W View FIGURE 4 )); in S. federicoescobari its excavation is deeper than wide in lateral view ( Fig. 4I View FIGURE 4 )).

Description. Body. Oval-elongate. Surface opaque, completely microgranulate. Color. Head and elytra dark brown. Other parts yellow or light brown. Length. 7.8–9.5 mm. Thorax. Anterior angles of pronotum approximately 85°. Lateral margin regularly curved outward, not forming an angle at the middle portion. Elytra. Striae thin and shiny, weakly impressed, punctures inconspicuous. Eighth stria with a thin carina at the anterior portion. Aedeagus. Parameres strongly asymmetrical ( Fig. 4R View FIGURE 4 ). Dorsal margin of right paramere curved inward, apex obliquely truncate. Ventral margin of right paramere substraight, with a rounded excavation at the basal portion. Dorsal margin of left paramere curved inward from the basal to medial portions, apical portion with a short and pointed projection. Ventral margin of left paramere substraight, with a bilobate excavation at the basal portion, apical portion pointed. SRP circular, with rounded handle-shaped extension ( Fig. 5Q View FIGURE 5 ). FLP short, comma-shaped, with three sets of bristles right beside it ( Fig. 5Q View FIGURE 5 ). A+SA with two superposed and elongate sclerites ( Fig. 5Q View FIGURE 5 ).

Type material. Holotype. BRAZIL: PARÁ: Pau d’Arco, Fazenda Marajoara (50°16’W, 7°50’S), 12.x.1998, P.Y. Scheffler— 1♂ ( CEMT) GoogleMaps . Paratypes [12 males, 9 females]. BRAZIL: PARÁ: Redenção, Pinkaiti Aik (52°02’W, 7°46’S), vi.1999, P.Y. Scheffler— 1♀ ( CEMT); same data but x.1999 — 1♂ ( CEMT); GoogleMaps Santarém, Reserva Tapajós, Amazon rainforest (55°39’842” W, 2°45’463”S), 25.xii.2008, pitfall, R.Andrade— 1♀ ( CEMT); GoogleMaps Curionópolis , Serra Leste (49°38’14”W, 5°58’27”S), 26.i.2016, pitfall, T. Portella— 4♂ ( CEMT); GoogleMaps Tucuruí (3°46’26’’S, 49°41’19’’W), i.1979, Alvarenga— 2♂ 1♀ ( CMNC). TOCANTINS, Araguaína , 22.xi.2014, Mata , Marta , & Sara— 1♂ (MZUF- PA) GoogleMaps ; Araguaína, viii.2014, M.L.S. Rippel— 1♂ ( MZUFPA); Araguaína, Universidade Federal do Tocantins-Escola de Medicina Veterinária e Zoologia, Mata de Galeria, Corrego da vaca, 16.iii.2017, pitfall 01, D. Moura & S. Dan-tas— 1♂ 2♀ ( MZUFPA); Araguaína, área aberta, 23.iii.2017, pitfall 04, D. Moura & S. Dantas— 2♀ ( MZUFPA); Araguaína, Universidade Federal Tocantins-Escola de Medicina Veterinária e Zoologia, 9.iv.2016, pitfall, S. Dan-tas— 2♂ 2♀ ( MZUFPA) .

Distribution. Known from Brazil (Pará and Tocantins) ( Fig. 6D View FIGURE 6 ). Endemism areas: Chacoan sub-region: South-eastern Amazonian dominion: Xingu-Tapajós province (see Morrone 2014; fig. 12).