Eumicrotremus gyrinops ( Garman 1892 )

Stevenson, Duane E., Mecklenburg, Catherine W. & Kai, Yoshiaki, 2017, Taxonomic clarification of the Eumicrotremus asperrimus species complex (Teleostei: Cyclopteridae) in the eastern North Pacific, Zootaxa 4294 (4), pp. 419-435: 430-434

publication ID

https://doi.org/10.11646/zootaxa.4294.4.2

publication LSID

lsid:zoobank.org:pub:151ACA96-34F5-4248-B795-1D9DB5F079FA

persistent identifier

http://treatment.plazi.org/id/9B6B4902-E176-C304-97C8-FE35D92CFE37

treatment provided by

Plazi

scientific name

Eumicrotremus gyrinops ( Garman 1892 )
status

 

Eumicrotremus gyrinops ( Garman 1892) 

Alaskan Lumpsucker

Figs. 3View FIGURE 3, 5View FIGURE 5; Table 1

Cyclopteroides gyrinops Garman 1892:37  . [Type locality: St. Paul Island , Alaska. Holotype: MCZAbout MCZ 16026View Materials.] 

Lethotremus muticus Gilbert 1896:449  . [Type locality: Albatross station 2844, near Unimak Pass, 54 fathoms. Lectotype: USNMAbout USNM 53806View Materials; Paralectotypes: CAS-SUAbout CAS-SU 774View Materials, CAS-SUAbout CAS-SU 3093View Materials, USNMAbout USNM 48614View Materials, USNMAbout USNM 59376View Materials.] 

Eumicrotremus phrynoides Gilbert & Burke 1912:69  . [Type locality: Albatross station 4779, Petrel Bank, Bering Sea, 54–56 fathoms. Holotype: USNMAbout USNM 74378View Materials.] 

Cyclopteropsis phrynoides: Soldatov & Lindberg 1930:325  . [New combination].

Cyclopterocottus phrynoides: Popov 1930:74  . [New combination].

Eumicrotremus gyrinops: Lindberg & Legeza, 1955:423  . [New combination].

Eumicrotremus muticus: Oku et al., 2017:55  . [New combination].

Holotype. MCZAbout MCZ 16026, 38.5 mm, St. Paul Island , Alaska, 56.9°N, 170.4°W.GoogleMaps 

Diagnosis. A species of Eumicrotremus  distinguished from all other ENPAbout ENP species of Eumicrotremus  by the following combination of characters: tubercles, when present, irregularly arranged in interorbital space (vs. four distinct interorbital rows in E. orbis  ); enlarged pair of tubercles absent at origin of second dorsal and anal fin (vs. present in E. andriashevi  ); thick dermal papillae absent (present in E. barbatus  ). Eumicrotremus gyrinops  is distinguished from members of the WNP E. asperrimus  species complex (including E. asperrimus  , Cyclopteropsis bergi  , and C. lindbergi  ) by higher modal counts of dorsal-fin spines (usually VII vs. usually VI) and lower modal counts of pectoral-fin rays (24–28 vs. 25–30).

Counts and proportions. First dorsal-fin spines VI –VIII; second dorsal-fin rays 10–12 (10.6); anal-fin rays 9– 12 (10.1); pectoral-fin rays 24–28 (26.1); vertebrae 10–12 + 15–19 = 26–30 (27.9). Following proportions as percent SL: head length 28.9–45.1 (36.9); body depth 28.7–57.7 (45.6); first dorsal-fin base 16.1–29.9 (23.5); second dorsal-fin base 15.4–26.2 (21.5); anal-fin base 13.7–25.4 (18.0); pectoral-fin base 24.8–35.8 (29.3); interdorsal distance 2.1–7–19.7 (7.9); predorsal length 29.1–47.2 (38.9); snout to vent 57.3–76.6 (64.6); vent to anal-fin origin 9.7–23.6 (16.3); caudal peduncle length 8.5–14.7 (11.6); caudal peduncle depth 8.0–13.0 (9.9); pectoral-fin length 15.6–26.8 (21.3); snout to disc 17.5–35.1 (24.4); disc length (outer) 20.2–30.3 (23.7); disc length (inner) 9.9–16.2 (12.1); disc width (outer) 18.5–29.3 (24.1); disc width (inner) 9.7–17.0 (13.3); disc to vent (outer) 8.5–31.1 (21.1); disc to vent (inner) 11.6–35.6 (25.6); caudal-fin length 22.5–31.9 (26.7). Following proportions as percent HL: orbit length 31.5–50.8 (40.4); snout length 7.4–25 (15.9); postorbital head length 32.7– 54.1 (41.2); interorbital width 40.9–78.5 (55.4); mouth width 50.4–93.9 (73.2); anterior internasal width 20.4–36.1 (28.9).

Description. Head and body short, firm, and globose; snout short; cheeks tumid. Interorbital space broad, slightly concave. Eye large, placed high on head, entering dorsal profile. Nostrils with tubes, the anterior short and wide, posterior relatively narrow, approximately same length as anterior. Mouth wide, slightly oblique, posterior margin of maxilla terminating anterior to margin of orbit. Teeth in jaws blunt, conical, arranged in three to five distinct diagonal rows on both premaxilla and dentary, approximately 25–35 teeth in each jaw. Upper and lower pharyngeal teeth present, similar in size and shape to jaw teeth.

Cephalic sensory system consisting of supraorbital, interorbital, postorbital, suborbital, and operculomandibular canals. Supraorbital canal opening through two pores, one anterior to nostrils and one medial to internarial space. Interorbital canal opening through single median pore. Postorbital canal opening anteriorly through a supraorbital pore and posteriorly through a postbranchial pore. Suborbital canal with two pores, anteriormost pore ventral to orbit, posterior pore posteroventral to orbit. Operculomandibular canal with single opercular pore on cheek and four mandibular pores closely spaced along lower jaw. All sensory pores opening through short thick tubes.

Development and distribution of bony tubercles on head and body highly variable. In holotype (38.5 mm SL), tubercles small, low, rounded, embedded in skin, each with several prominent spines; tubercles few and sparsely spaced, forming an indistinct row on either side of spinous dorsal fin, several additional tubercles dispersed along anterior portion of body; head naked. Tubercles generally smaller and fewer in smaller specimens and in males; larger males (45–55 mm SL) with tubercles completely covered by skin, producing wart-like protrusions on body; males and smaller females often without tubercles on head or body; all specimens examined over 58 mm SL (n = 13) completely covered with large tubercles, and all female.

First dorsal fin high, usually not covered with thick skin or tubercles in smaller specimens (including holotype); becoming increasingly embedded in skin and covered with spiny tubercles in larger specimens, particularly in females; fin membranes moderately to heavily pigmented. Second dorsal and anal fins directly opposite each other, nearly equal in length and height, usually without bony tubercles or fleshy papillae; fin membranes translucent, without pigment. Caudal fin truncate; fin membranes translucent, without pigment; rays in some specimens with several faint dark blotches suggesting bands running perpendicular to body axis. Pectoral fins translucent, their bases densely covered with small melanophores; larger specimens usually with several rounded tubercles covering pectoral base. Pelvic disc large, ovate, slightly constricted anteriorly; its length usually slightly greater than its width. Anus approximately midway between posterior margin of pelvic disc and origin of anal fin.

Coloration. Holotype with small melanophores relatively evenly spaced over entire head and body, without conspicuous bands, stripes, or blotches. Additional material in life yellowish-brown to medium brown, generally darker on head and dorsum, often with dark brown to black blotches and vermiculations, particularly on dorsal surface of head and body; becoming lighter posteriorly and generally white or nearly white on ventral surface. Some degree of faint banding usually present on pectoral and all median fins.

Distribution. Specimens examined are from the eastern Bering Sea, Aleutian Islands, and western and central Gulf of Alaska ( Fig. 6View FIGURE 6).

Remarks. The holotype of E. gyrinops  ( MCZAbout MCZ 16026View Materials) is in poor condition. It has apparently been desiccated and rehydrated at least once. Thus, the tubes associated with the mandibular pores, and to a lesser extent the other pores as well, appear longer and more prominent than in most other specimens. In addition, its fins are badly damaged, and its internal organs have been removed through a very large incision on the left side of the body.

Material examined. Eumicrotremus asperrimus  (18 specimens): Sea of Japan : FAKUAbout FAKU 131263 (GenBank ABView Materials 917623, 917693), 102 mm female, Kyoto, 36°N, 134.75°E, 320 m  ; FAKU 131983 (GenBank AB917614, 917684), 40 mm male, 35.881°N, 132.474°E, 240 m; FAKU 132544 (GenBank AB917600, 917670), 59 mm male, 37.758°N, 136.294°E, 342 m; FAKU 132564 (GenBank AB917601, 917671), 58 mm male, 38.026°N, 136.916°E, 330 m; FAKUAbout FAKU 132598 (GenBank AB917621View Materials, 917691), 34 mm male, Ishikawa, 37.7°N, 136.308°E, 260 mGoogleMaps  ; FAKUAbout FAKU 133204 (GenBank ABView Materials 917615, 917685), 61 mm male, Kyoto, 38.03°N, 135.413°E, 300 m. Sea of Okhotsk : FAKUAbout FAKU 133231View Materials (GenBank AB917616View Materials, 917686), 98 mm female, 45.062°N, 143.068°EGoogleMaps  ; FAKU 133235 (GenBank AB917620, 917690), 52 mm, 44.99°N, 144.115°E, 220 m; FAKU 133279 (GenBank AB917604, 917674), 56 mm female, 44.382°N, 144.341°E, 350 m; FAKU 133280 (GenBank AB917605, 917675), 54 mm female, 44.382°N, 144.341°E, 350 m; FAKU 133281 (GenBank AB917606, 917676), 59 mm female, 44.382°N, 144.342°E; FAKU 133282 (GenBank AB917607, 917677), 60 mm male, 44.382°N, 144.342°E; FAKU 133283 (GenBank AB917608, 917678), 60 mm female, 44.382°N, 144.342°E; FAKU 133284 (GenBank AB917609, 917679), 49 mm male, 44.382°N, 144.341°E, 350 m; FAKU 133285 (GenBank AB917610, 917680), 91 mm female, 44.382°N, 144.342°E; FAKU 133286 (GenBank AB917611, 917681), 56 mm female, 44.382°N, 144.341°E, 350 m; FAKU 134958 (GenBank AB917639, 917709), 59 mm female, 44.78°N, 143.928°E, 170 m; FAKU 134985 (GenBank AB917643, 917713), 56 mm male, 45.46°N, 142.967°E, 117 m.

Eumicrotremus awae  : FAKU 134183, 18 mm, Japan, Sagami Bay; FAKU 134725, 18 mm, Japan, Sagami Bay; FAKU 135824, 24 mm, Japan, Sagami Bay; UW 42938, 16 mm, Japan, Sagami Bay, 35.15°N, 139.17°E.

Eumicrotremus bergi  (2 specimens): FAKUAbout FAKU 132552View Materials (GenBank AB917597View Materials, 917667), 59 mm male, Sea of Japan , 38.031°N, 136.778°E, 419 m; FAKUAbout FAKU 132557View Materials (GenBank AB917596View Materials, 917666), 62 mm male, Sea of Japan, 37.959°N, 137.31°E, 320 m.GoogleMaps 

Eumicrotremus gyrinops  (235 specimens): Type material: MCZAbout MCZ 16026View Materials, holotype of Cyclopteroides gyrinops  , 38.5 mm, St. Paul Island , Alaska ; USNMAbout USNM 74378View Materials, holotype of Eumicrotremus phrynoides, Albatross  station 4779, Petrel Bank , Bering Sea , 54–56 fm; USNMAbout USNM 53806, lectotype of Lethotremus muticus, Albatross  station 2844, near Unimak Pass , 54 fm; CAS-SUAbout CAS-SU 774, paralectotype of Lethotremus muticus, Albatross  station 2844, near Unimak Pass, 54 fm; CAS-SUAbout CAS-SU 3093, USNMAbout USNM 48614, USNMAbout USNM 59376, paralectotypes of Lethotremus muticus, Albatross  station 3223, Unimak Pass, 39 fm. Bering Sea (31 specimens): UW 28373, 85 mm female, 57°0'N, 171°25.02'W, 59 fm; UW 28384, 69 mm female, 60°59.1'N, 177°19.02'W, 130 mGoogleMaps  ; UW 28390, 65 mm female, 60°28.14'N, 177°W, 143 m; UW 119071, 16(29–47 mm males, 27–40 mm females), 56°53'N, 169°39'W, 75 m; UW 119538 (GenBank AB917661, 917731), 92 mm female, 57°59.23'N, 170°20.09'W, 74 m; UW 150056, 82 mm female, 59°19.98'N, 172°28.26'W, 87 m; UW 150057, 51 mm male, 56°50.08'N, 170°26.76'W, 98 m; UW 150071 (GenBank AB917626, 917696), 75 mm female, 54°50.25'N, 169°57.12'W, 71 m; UW 150072, 88 mm female, 60°20.28'N, 174°3.36'W, 90 m; UW 151282, 3(41 mm male, 33–77 mm females), 56°49.57'N, 170°28.07'W, 101 m; UW 151285, 80 mm female, 58°20.58'N, 170°22.32'W, 74 m; UW 151415, 46 mm male, 56°40.32'N, 170°4.44'W, 95 m; UW 152368, 36 mm male, 60°8.88'N, 172°57.12'W, 59 m; UW 155892, 81 mm female, 56°35.22'N, 170°38.22'W, 108 m.

Aleutian Islands (160 specimens): UW 22199, 69 mm, 53°43.98'N, 164°49.98'W, 209 mGoogleMaps  ; UW 28389, 42 mm, 54°24'N, 165°24'W; UW 46659, 32 mm, 51°54.6'N, 176°52.2'W, 211 m; UW 46660, 45 mm, 52°51.53'N, 172°27.52'E, 146 m; UW 46662, 45 mm, 53°1.68'N, 173°12.36'E, 133 m; UW 46664, 5(43–51 mm females), 52°11.04'N, 179°39.84'W, 115 m; UW 46665, 5(44–49 mm females), 51°58.98'N, 179°23.52'E, 139 m; UW 49401, 2(40 mm male, 45 mm female), 52°7.8'N, 179°54.6'W, 112 m; UW 49409, 41 mm male, 52°4.92'N, 179°21.54'E, 306 m; UW 49412, 45 mm male, 52°25.86'N, 179°56.52'E, 163 m; UW 49418, 49 mm male, 52°31.27'N, 174°26.96'E, 182 m; UW 49423, 50 mm female, 52°2.82'N, 179°25.26'E, 143 m; UW 49436 ( GenBank AB917625View Materials, 917629, 917695, 917699), 2(36–38 mm, males), 52°08.14'N, 179°53.41'W, 121 mGoogleMaps  ; UW 49467, 34 mm male, 52°29.96'N, 170°40.65'W, 208 m; UW 49470, 4(37–52 mm females, 36 mm male), 52°10.86'N, 179°37.02'E, 124 m; UW 49471, 4(41–44 mm males), 52°22.62'N, 179°38.82'E, 245 m; UW 49474, 18(34–38 mm males, 30–40 mm females), 51°39.23'N, 176°22.56'W, 143 m; UW 49504, 61 mm female, 52°2.34'N, 179°24.96'E, 145 m; UW 49577, 5(40–42 mm females), 52°59.42'N, 172°21.95'E, 146 m; UW 49580, 49 mm female, 52°28.15'N, 173°11.34'E, 116 m; UW 49587, 45 mm female, 52°35.6'N, 172°55.38'E, 154 m; UW 111260, 39 mm male, 52°27.91'N, 174°12.6'E, 110 m; UW 111265, 35 mm female, 52°23.82'N, 173°58.72'E, 112 m; UW 111267, 2(34– 41 mm females), 51°39.23'N, 176°22.2'W, 145 m; UW 111280, 52 mm female, 52°52.7'N, 175°12.5'W, 151 m; UW 111288, 62 mm female, 52°25.27'N, 170°16.74'W, 211 m; UW 111300, 47 mm female, 52°21.71'N, 171°16.14'W, 134 m; UW 111309, 40 mm male, 52°21.96'N, 179°54.84'W, 168 m; UW 111310, 35 mm female, 52°29.56'N, 174°22.08'W, 177 m; UW 111311, 7(32–42 mm males, 38–51 mm females), 52°57.72'N, 170°23.82'W, 221 m; UW 111314, 2(49–50 mm females), 52°18.38'N, 175°48.91'E, 235 m; UW 111335, 35 mm male, 52°30.19'N, 174°15.22'W, 102 m; UW 111451, 2(36–44 mm males), 52°29.46'N, 170°8.82'W, 183 m; UW 112040, 35 mm male, 52°22.97'N, 174°8.62'E, 148 m; UW 112041, 41 mm female, 52°26.26'N, 174°17.57'E, 220 m; UW 112272View Materials ( GenBank AB917663View Materials, 917664, 917733, 917734), 2(32 mm male, 45 mm female), 52°31.20'N, 170°39.25'W, 124 mGoogleMaps  ; UW 117593, 43 mm female, 52°39.45'N, 173°50.48'E, 79 m; UW 151458, 3(35–38 mm males, 33 mm female), 51°2.50'N, 172°15.1'W, 162 m; UW 151803View Materials ( GenBank AB917644View Materials, 917714), 36 mm male, 51°53.15'N, 173°55.29'W, 136 mGoogleMaps  ; UW 151804View Materials ( GenBank AB917645View Materials, 917715), 45 mm female, 52°02.56'N, 179°48.44'W, 250 mGoogleMaps  ; UW 151810View Materials ( GenBank AB917646View Materials, 917716), 43 mm female, 52°49.40'N, 173°41.17'W, 109 mGoogleMaps  ; UW 151811View Materials ( GenBank AB917647View Materials, 917717), 44 mm female, 52°24.33'N, 170°16.57'W, 213 mGoogleMaps  ; UW 151814View Materials ( GenBank AB917648View Materials, 917718), 46 mm male, 52°49.42'N, 170°27.32'W, 99 mGoogleMaps  ; UW 152131View Materials ( GenBank LC209624View Materials, 209640), 49 mm female, 52°35.48'N, 172°55.48'E, 145 mGoogleMaps  ; UW 152132View Materials ( GenBank LC209625View Materials, 209641), 40 mm male, 52°28.32'N, 173°11.4'EGoogleMaps  ; UW 152138, 2(31 mm male, 33 mm female), 54°22.2'N, 165°32.94'W, 83 m; UW 152355, 3(39–60 mm male, 41 mm female), 52°55.94'N, 170°24.61'W, 222 m; UW 152358View Materials ( GenBank LC209626View Materials, 209642), 35 mm female, 51°45.04'N, 177°51.7'E, 157 mGoogleMaps  ; UW 152359View Materials ( GenBank LC209627View Materials, 209643), 3(38 mm male, 37–41 mm females), 51°54.64'N, 173°47.03'W, 118 mGoogleMaps  ; UW 152360View Materials ( GenBank LC209628View Materials, 209644), 33 mm male, 51°39.08'N, 178°20.06'E, 206 mGoogleMaps  ; UW 152361, 4(36–42 mm males, 33 mm female), 52°29.62'N, 170°23.63'W, 216 m; UW 152362View Materials ( GenBank LC209629View Materials, 209645), 32 mm male, 52°2.16'N, 179°24.84'E, 150 mGoogleMaps  ; UW 152363View Materials ( GenBank LC209630View Materials, 209646), 29 mm female, 51°36.96'N, 178°11.28'W, 134 mGoogleMaps  ; UW 152364, 38 mm female, 52°17.28'N, 170°47.58'W, 226 m; UW 152365View Materials ( GenBank LC209631View Materials, 209647), 35 mm male, 52°49.68'N, 173°41.28'E, 115 mGoogleMaps  ; UW 152366View Materials ( GenBank LC209632View Materials, 209648), 36 mm male, 51°52.92'N, 173°55.38'WGoogleMaps  ; UW 152367View Materials ( GenBank LC209633View Materials, 209634, 209649, 209650), 2(37–41 mm males), 52°52.02'N, 171°18.06'W, 204 mGoogleMaps  ; UW 152369View Materials ( GenBank LC209635View Materials, 209651), 33 mm female, 51°47.28'N, 177°43.56'E, 155 mGoogleMaps  ; UW 152370View Materials ( GenBank LC209636View Materials, 209652), 3(40–41 mm females), 51°37.98'N, 177°9.96'W, 172 mGoogleMaps  ; UW 152371View Materials ( GenBank LC209637View Materials, 209653), 2(35–37 mm males), 52°26.88'N, 174°18.66'E, 228 mGoogleMaps  ; UW 152428, 53 mm female, 52°52.28'N, 171°19.06'W, 202 m; UW 152431 (LC209833, 209845), 40 mm female, 52°22.92'N, 174°8.46'E, 155 m; UW 152432, 43 mm female, 52°26.88'N, 174°18.66'E, 228 m; UW 152433View Materials ( GenBank LC209638View Materials, 209639, 209654, 209655), 2(35–42 mm males), 52°58.62'N, 170°3.72'W, 122 mGoogleMaps  ; UW 152434 (LC209834, 209846), 41 mm male, 52°28.33'N, 173°11.37'E; UW 152438, 2(27–43 mm males), 52°28.59'N, 170°8.7'W, 183 m; UW 152444, 39 mm male, 54°20.65'N, 165°32.94'W, 104 m; UW 152460, 2(38 mm male, 31 mm female), 52°23.18'N, 174°W, 85 m; UW 152466, 2(36 mm male, 46 mm female), 52°48.65'N, 171°31.38'W, 201 m; UW 152469 (LC209835–209843, 209847–209855), 9(37–48 mm females), 52°48.95'N, 173°44.35'E, 111 m; UW 152490 (LC209844, 209856), 42 mm female, 52°24.22'N, 174°15.53'E, 255 m; UW 154964, 9(41 mm male, 36–57 mm females), 52°10.84'N, 179°36.97'E, 127 m; UW 156510, 4(32–38 mm females), 51°47.46'N, 177°40.74'E, 171 m; UW 156512, 37 mm female, 52°28.5'N, 170°44.94'W, 260 m; UW 156511, 2(41–42 mm females), 52°35.28'N, 172°55.72'E, 127 m.

Gulf of Alaska (42 specimens): UW 28386, 2(52–54 mm), 56°N, 157°W, 50 fm; UW 47916, 38 mm female, 58°7.47'N, 151°30.6'W, 157 mGoogleMaps  ; UW 47923, 9(34–39 mm females, 37 mm male), 58°56.46'N, 150°19.02'W, 146 m; UW 47935, 2(39 mm female, 39 mm male), 56°43'N, 152°36.72'W, 97 m; UW 47948, 3(32 mm female, 37–40 mm males), 54°53.85'N, 158°42.48'W, 98 m; UW 116417, 40 mm female, 58°46.32'N, 150°23.58'W, 151 m; UW 117047, 4(34–38 mm males, 36–38 mm females), 54°44.1'N, 158°37.26'W, 104 m; UW 117609View Materials ( GenBank AB917634View Materials, 917704), 3(41–43 mm females, 38 mm male), 58°15.17'N, 151°24.51'W, 157 mGoogleMaps  ; UW 119222View Materials ( GenBank AB917627View Materials, 917628, 917697, 917698), 2(32–38 mm females), 57°36.33'N, 150°20.16'W, 130 mGoogleMaps  ; UW 119548, 39 mm male, 55°16.88'N, 157°6.48'W, 94 m; UW 119596, 40 mm male, 57°53.28'N, 150°16.47'W, 158 m; UW 119690, 4(38–41 mm females), 58°53.03'N, 150°52.08'W, 164 m; UW 119706, 60 mm female, 58°54.29'N, 150°36.3'W, 119 m; UW 119779View Materials ( GenBank AB917631View Materials, 917701), 35 mm male, 58°38.59'N, 150°08.45'W, 121 mGoogleMaps  ; UW 119782View Materials ( GenBank AB917632View Materials, 917702), 39 mm male, 58°30.48'N, 150°17.52'W, 90 mGoogleMaps  ; UW 151260View Materials ( GenBank AB917657View Materials, 917658, 917659, 917727, 917728, 917729), 5(31–37 mm males, 35 mm female), 58°54'N, 151°24.96'W, 153 mGoogleMaps  ; UW 154590, 39 mm male, 55°29.21'N, 157°2.82'W, 90 m.

Eumicrotremus lindbergi  (4 specimens): FAKUAbout FAKU 132535View Materials ( GenBank AB917599View Materials, 917669), 55 mm male, Sea of Japan, 36.845°N, 136.317°E, 261 mGoogleMaps  ; FAKUAbout FAKU 132563View Materials ( GenBank AB917598View Materials, 917668), 59 mm male, Sea of Japan, 38.026°N, 136.916°E, 330 mGoogleMaps  ; FAKUAbout FAKU 134222View Materials (GenBank AB917624View Materials, 917694), 53 mm male, Sea of Japan, Shimane, west of Oki I.  ; FAKUAbout FAKU 134862View Materials ( GenBank AB917638View Materials, 917708), 54 mm male, Sea of Japan, Shimane, 35.6°N, 131.033°E, 180 m.GoogleMaps 

Eumicrotremus orbis  : UW 119821View Materials (GenBank AB917655View Materials, 917725), 2(17 mm male, 31 mm female), Salish Sea , San Juan Island, Jackson Beach, 48°31.2'N, 123°W, 2 m. 

MCZ

Museum of Comparative Zoology

USNM

Smithsonian Institution, National Museum of Natural History

CAS-SU

California Academy of Sciences, Stanford University Collection

ENP

Everglades National Park

FAKU

Kyoto University

Kingdom

Animalia

Phylum

Chordata

Class

Actinopterygii

Order

Scorpaeniformes

Family

Cyclopteridae

Genus

Eumicrotremus

Loc

Eumicrotremus gyrinops ( Garman 1892 )

Stevenson, Duane E., Mecklenburg, Catherine W. & Kai, Yoshiaki 2017

2017
Loc

Cyclopteroides gyrinops

Garman 1892: 37

Loc

Lethotremus muticus

Gilbert 1896: 449

Loc

Eumicrotremus phrynoides

Gilbert 1912: 69

Loc

Cyclopteropsis phrynoides:

Soldatov 1930: 325

Loc

Cyclopterocottus phrynoides:

Popov 1930: 74

Loc

Eumicrotremus gyrinops:

Lindberg 1955: 423

Loc

Eumicrotremus muticus: Oku et al., 2017 :55

Oku 2017: 55