Manota anceps, Hippa, Heikki & Ševċík, Jan, 2010

Hippa, Heikki & Ševċík, Jan, 2010, Notes on Oriental and Australasian Manotinae (Diptera, Mycetophilidae), with the description of thirteen new species, Zootaxa 2333, pp. 1-25 : 4-6

Manota anceps sp. n.

Figs 3 A, B, C

Male. Colour. Head pale brown, frons and vertex darker brown. Antenna unicolorous pale brown. Mouthparts pale yellowish-brown. Thorax unicolorous pale brown. Legs unicolorous pale yellowish. Wing unicolorous yellowish; haltere yellowish-brown with darker brown knob. Abdominal tergites brownish, sternites paler yellowish-brown, sternites 7 and 8 almost as dark as tergites. All setosity yellowish or brownish, thicker setae seeming darker than finer setae and trichia. Head. Antennal flagellomere 4, Fig. 3 A. Palpomere 3 of maxillary palpus with apicomesial extension, with 5 apically expanded curved sensilla; palpomere 4 with parasegment; palpomere 5 1.4 times as long as palpomere 4. Nine strong postocular setae. Thorax.

Anepisternum setose, with 41 setae; anterior basalare non-setose; preepisternum 2 setose, with ca. 20 setae; laterotergite non-setose; episternum 3 setose, with ca. 15 setae. Wing. Similar to Fig. 1 A; wing length 1.6 mm. Hypopygium, Figs 3 B, C. Sternite 9 about two thirds of ventral length of gonocoxa, with sharply delimited convex sides, posterior margin transverse, anterior margin deeply incised, setae similar to the adjacent ventral setae of gonocoxa. Ventral mesial margin of gonocoxa simple, roundly curved, with a discontinuity at the middle. Parastylar lobe oblique, slightly sickle-shaped, with two setae antero-mesially. Paraapodemal lobe well exposed in ventral view. Dorsal mesial margin of gonocoxa simple, convex, subapically with two stronger setae and apically with two strong setae arising from large basal bodies, united basally. Two juxtagonostylar setae present, both long almost straight megasetae, arising from a common basal body nearly half the length of the megasetae. Gonostylus simple, obovate in outline, ventrally with moderately long setosity, at mesial margin forming a fringe of more tightly placed setae, dorsally non-setose. Aedeagus short subtriangular with weak lateral shoulders, the apex curved ventrad. Hypoproct posteriorly extending to the level of base of gonostylus, ventrally with a mesial row of 5 setae (sternite 10) on each half. Cerci mesially separate.

Female unknown.

Discussion. M. anceps belongs to a common form of Oriental Manota by having a setose anepisternum and preepisternum 2, a non-setose anterior basalare and laterotergite, a short vein R1, a laterally free tergite 9, a sickle-shaped parastylar lobe, the two subequal juxtagonostylar megasetae arising from a long common basal body and a rather unmodified gonostylus. Among these species M. anceps resembles those which have one or more apicomesial setae arising from a long basal body on the dorsal side of the gonocoxa, strongly resembling the juxtagonostylar setae. These species are M. duplex Hippa (Malay Peninsula), M. pellii Hippa ( Thailand) , M. perpusilla Hippa (Malay Peninsula), and M. vesicaria Hippa ( Thailand) . M. pellii , M. perpusilla and M. vesicaria have one such seta strong and M. duplex 2–3 fine setae. M. anceps is the only species with two setae arising from separate basal bodies which unite at the base. M. perpusilla and M. duplex differ further from the species mentioned above e.g. by having a conspicuously longer and narrower gonostylus. M. vesicaria differs by having the paraapodemal lobe inflated and the setae of the hypoproct all placed laterally and M. pellii by having three setae on the parastylar lobe.

Etymology. The name is derived from the Latin anceps , two-headed, double. It refers to the two-headed lobe dorsally at the apex of the gonocoxa, resembling a duplicate of the juxtagonostylar megasetae.

Types. Holotype. Male, INDONESIA, Sumatra, Aceh, Gunung Leuser National Park, Ketambe res. st., 3o41’N, 97o39’E, young forest, light gap, 350 m, 1–31.i.1990, Malaise trap, D.C. Darling, IIS 900001 (in ROM).

Figures

FIGURE 1. Wing, dorsal view (A, B, C, E) and basal part of wing, ventral view (D). A. Manota bruneiensis sp. n. (paratype) B. M. subforceps sp. n. (holotype). C. M. sinepollex sp. n. (holotype). D, E. Eumanota vilkamaai sp. n. (holotype). Scale 1.0 mm.

FIGURE 3. Manota anceps sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. Scale 0.10 mm.