Tofanella Bandel, 1995

Gründel, Joachim & Nützel, Alexander, 2013, Evolution and classification of Mesozoic mathildoid gastropods, Acta Palaeontologica Polonica 58 (4), pp. 803-826 : 815

Genus Tofanella Bandel, 1995

Figs. 8B , 9A, B .

Type species: Turritella decussata von Münster, 1841 , original designation; Late Triassic , North Italy .

Included species: Turritella decussata Münster, 1841 , Late Triassic; Tofanella cancellata Bandel, 1995 , Late Triassic;? Tricarilda octoangulata Gründel, 2006 , Bathonian; several undescribed species from the Pliensbachian of England (JG, own observation).

Diagnosis.—“The turriculate shell has a major keel on the first whorls of the teleoconch, which disappears on later whorls as they become almost flat. The spiral sculpture is crossed by few collabral elements. The protoconch has a smooth surface, and the embryonic shell is immersed in its apex. In the larval whorls the sinistral coiling changes into dextral coiling before onset of the teleoconch. With transition from larval to adult shell sculpture and whorl shape change drastically” ( Bandel 1995: 21).

Stratigraphic and geographic range.—Late Triassic to Pliensbachian; England,? France, North Italy.

Figures

Fig. 8. Coaxial heterostrophic protoconchs of Mathilda and Tofanella sensu Bandel (1995) do not show principal differences. A. Mathilda bolina (von Münster, 1841) sensu Bandel (1995), Alpe di Specie, N Italy, Late Triassic; from Bandel (1995: pl. 2: 2). B. Tofanella cancellata Bandel, 1995, Cassian Formation, Campo, Dolomites, N Italy, Late Triassic; from Bandel (1995: pl. 11: 8).

Fig. 9. Type species and examples for Triassic and Jurassic genera of the family Tofanellidae. A, B. Tofanella laevigata (von Münster, 1841), Cassian Formation, Alpe di Specie, Dolomites, N Italy, Late Triassic A. From Bandel (1995: pl. 10: 1). B. From Bandel (1995: pl. 10: 2). C. Cristalloella (Cristalloella) cassiana Bandel, 1995, Cassian Formation, Campo, Dolomites, N Italy, Late Triassic; from Bandel (1995: pl. 10: 9). D. Cristalloella (Wonwalica) minuta Schröder, 1995, Wąwał, Poland, Valanginian; from Schröder (1995: pl. 10: 1). E. Neodonaldina elongata (Zardini, 1978), Cassian Formation, Alpe di Specie, Dolomites, N Italy, Late Triassic. Protoconch (E1) from Bandel (1996: fig. 5d), whole specimen (E2) from Bandel (1996: fig. 6a). F. Graphis rotundata (Gründel, 1998), borehole Heringsdorf 4/63, Germany, middle Callovian; from Gründel (1998: pl. 1: 5). G. Graphis albida (Kanmacher, 1798), Pliocene, Nice/France; from Bandel (2005: fig. 64). H. Camponaxis lateplicata (Klipstein, 1843), Cassian Formation, Misurina, Dolomites, N Italy, Late Triassic; from Bandel (1995: pl. 14: 3). I. Usedomella laevigatoidea (Gründel, 1993), borehole Kłęby 1/37 (formerly Klemmen), Poland, Callovian; from Gründel (1998: pl. 2: 19). J. Urlocella minuera Gründel, 1998, bore Kłęby 1/37, late Bathonian; from Gründel (1998: pl. 1: 11).