Blastobasis echus Adamski

Adamski, David, 2013, Review of the Blastobasinae of Costa Rica (Lepidoptera: Gelechioidea: Blastobasidae), Zootaxa 3618 (1), pp. 1-223 : 50-53

Blastobasis echus Adamski , new species

( Figs. 17 , 99–100 , 293 , 336 , Map 14)

Diagnosis.— Blastobasis echus is similar to B. usurae in facies but differs from the latter by having a wider anal opening; a more elongate pigmented part of the gnathos; sockets of tergal setae extending beyond the midlength of the tegumen; a wider proximal flange of the dorsal part of the valva; a larger phallus; and an apex of the anellus that is more emarginate mesially. B. echus also has and an anellus that is setose only along its lateral margins, a feature that is lacking in B. usurae .

Description.—Head: Scales on vertex and frontoclypeus brownish gray tipped with pale brownish gray. Outer and inner surfaces of labial palpus with brownish-gray scales tipped with pale brownish gray, and pale brownishgray scales along apical margins of segments 1–2. Antennal scape with brownish-gray scales tipped with pale brownish gray, pecten brown, flagellum dark brownish gray; first flagellomere in male dilated, inner surface of dilated part with 56 palmate sex scales (n=1), ( Fig. 293 ). Proboscis pale brownish gray.

Thorax: Tegula with brownish-gray scales tipped with pale brownish gray; mesonotum with basal 1/4 brownish gray, apical 3/4 with scales brownish gray tipped with pale brownish gray. Forewing ( Fig. 336 ): Length 4.1–8.5 mm (n = 140), brownish-gray scales tipped with pale brownish gray intermixed with brownish-gray scales and pale brownish-gray scales; basal 1/3 pale brownish gray intermixed with few brownish-gray scales except, costa brownish gray or base dark gradually brightening to 1/3; submedian fascia complete or incomplete; apical 2/ 3 dark basally, gradually brightening to apex; cell with three spots, one near middle, two on apical end along crossvein; marginal spots appear as short streaks on apical portions of radial and medial veins. Undersurface brown. Venation ( Fig. 17 ) with M3 and CuA1 arising from a common point on distoposterior part of cell; cubital veins divergent from bases, CuA1 slightly curved, CuA2 broadly curved. Hindwing: Translucent pale brown. Venation ( Fig. 17 ) with anterior part of cell closed, posterior part of cell open; cubitus 4-branched with all veins arising submarginally from cubitus.

MAP 14. Distribution of Blastobasis echus (●).

Abdomen: Male genitalia ( Figs. 99–100 ): Uncus slightly narrowed from base, broadly rounded apically, slightly downcurved, sparsely setose; shorter than width of anal opening. Dorsal strut of tegumen absent. Gnathos wide, confluent with tegumen, posteroventral margin slightly protracted mesially, bidentate; midwidth of pigmented part of gnathos narrower than midwidth of uncus. Sockets of tergal setae extending beyond midlength of tegumen. Valva divided; ventral part angled dorsolaterally from beyond base, narrowed apically, forming inwardly curved spinelike process; process setose on outer surface, planate on inner surface; ventral margin setose from about 1/5, upturned greatly beyond base, forming wide fold to near base of apical process; dorsal part with apical portion of costa extending dorsolaterally, forming setose digitate process; digitate process broadly curved inwardly; area beneath costa overlaid with densely packed microtrichiate membrane extending above and below phallus to opposite side; basal ridge of digitate process protracted ventrally fusing with dorsolateral ridge of proximal flange; ventral margin of proximal flange slightly cleft, extending laterally to a broadly rounded, apicoventral ridge; proximal flange bearing densely packed microtrichiae. Juxta bandlike. Vinculum semicircular. Phallus longer than valva, bulbous basally; sclerite of phallus shorter than valva, acutely curved near 2/3; anellus near parallelsided from base, apically notched mesially, setose along margin. Female Genitalia: Unknown.

Holotype, 3, “F[in]ca Cafrosa, Est[ación] Las Mellizas, P[arque] N[acional] Amistad, 1300 m, Prov[incia] Punt[arenas]”, “ COSTA RICA, M. Ramirez & G. Mora, Set [iembre] 1990, L-S-316100, 596100”, “INBio: COSTA RICA: CRI000, 666167” [barcode label], “INBio, 3 Genitalia Slide by D. Adamski, No. 3710” [yellow label].

Paratypes (139 3): 1 3, same data as for holotype except, “Set.”, “CRI000, 666148”, “Slide No. 3678”; 1 3, “Oct.”, “CRI000, 575391”, “Slide No. 3636”, “USNM 83790”; 2 3, “Nov.”, “CRI000, 521710”, “Slide No. 3698”, “USNM 83791”; “CRI000, 521859”, “Slide No. 3699”, “USNM 83792”; 1 3, “M. Ramirez, Mar., 1991”, “CRI000, 301699”, “Slide No. 3696”; 1 3, “Abr.”, “CRI000, 474616”, “Slide No. 3612”; 1 3, “M.M. Chavarria & G. Mora, Ene. 1991 ”, “CRI000,380751”, “Slide No. 3615”, “USNM 83793”; 13 3, “M. Ramirez, Abr. 1991 ”, “CRI000, 474700”, “Slide No. 3618”, “USNM 83794”; “CRI000, 474740”, Slide No. 3621” “CRI000, 474726”, “Slide No. 3624”, “USNM 83795”; “CRI000, 474809”, “Slide No. 3625”; “CRI000, 475010”, “Slide No. 3627”; “CRI000, 474760”, “Slide No. 3626”, “USNM 83796”; “CRI000, 474825”, “Slide No. 3655”; “CRI000, 474586”, “Slide No. 3681”; “CRI000, 474604”, “Slide No. 3658”, “USNM 83797”; “CRI000, 474869”, “Slide No. 3656”, “USNM 83798”; “CRI000, 474833”, “Slide No. 3682”; “CRI000, 475001”, “Slide No. 3688”; “CRI000, 474790”, “Slide No. 3609”; 3 3, “M. Ramirez, May. 1991 ”, “CRI000, 355324”, “Slide No. 3708”, “USNM 83799”; “CRI000, 355235”, “Slide No. 3657”; “CRI000, 355314”, “Slide No. 3670”, “USNM 83800”; 2 3, “G. Mora , May. 1991 ”, “CRI000, 320052”, “Slide No. 3717”, “USNM 83801”; “CRI000, 320086”, “Slide No. 3720”, “USNM 83802”; 10 3, “Est. Las Pailas, 800 m, P. N. Rincon de la Vieja, Prov. Guanacaste, COSTA RICA, 1 a 22 Jul. 1992, D. Garcia, L-N-306300, 388600”, “CRI000, 689983”, “Slide No. 3339”; “CRI000, 689963”, “Slide No. 3354”; “CRI000, 689978”, “Slide No. 3347”; “CRI000, 689981”, “Slide No. 3348”; “CRI000, 689956”, “Slide No. 3349”, “USNM 83803”; “CRI000, 689935”, “Slide No. 3351”, “USNM 83804”; “CRI000, 689979”, “Slide No. 3352”, “USNM 83805”; “CRI000, 689940”, “Slide No. 3355”; “CRI000, 689927”, “Slide No. 3358”; “CRI000, 689966”, “Slide No. 3359”, “USNM 83806”; 9 3, “C. Cano”, “CRI000, 718963”, “Slide No. 3341”; “CRI000, 718964”, “Slide No. 3330”; “CRI000, 719000”, “Slide No. 3328”; “CRI000, 718966”, “Slide No. 3336”; “CRI000, 719019”, “Slide No. 3334”; “CRI000, 718935”, “Slide No. 3333”, “USNM 83807”; “CRI000, 719017”, “Slide No. 3313”; “CRI000, 719004”, “Slide No. 3332”; “CRI000, 718933”, “Slide No. 3340”, “USNM 83808”; 18 3, “ 21–30 Nov. 1991 ”, “CRI000, 818562”, “Slide No. 3377”, “USNM 83809”; “CRI000, 818723”, “Slide No. 3378”; “CRI000, 818716”, “Slide No. 3379”, “USNM 83810”; “CRI000, 818801”, “Slide No. 3380”; “CRI000, 818797”, “Slide No. 3384”; “CRI000, 818613”, “Slide No. 3386”; “CRI000, 818629”, “Slide No. 3387”; “CRI000, 818594”, “Slide No. 3388”; CRI000, 818628”, “Slide No. 3389”; “CRI000, 818648”, “Slide No. 3390”; “CRI000, 818671”, “Slide No. 3391”; “CRI000, 828144”, “Slide No. 3318”, “USNM 83811”; “CRI000, 828085”, “Slide No. 3325”; “CRI000, 818621”, “Slide No. 3315”; “CRI000, 818778”, “Slide No. 3316”; “CRI000, 828122”, “Slide No. 3296”, “USNM 83812”; “CRI000, 818564”, “Slide No. 3431”; “CRI000, 818795”, “Slide No. 3430”; 2 3, “ 23 Set.-12 Oct., 1992, D. Garcia”, “CRI000, 863244”, “Slide No. 3293”; “CRI000, 920483”, “Slide No. 3292”; 3 3, “ 23 Oct.-12 Nov., 1992, C. Cano”, “CRI000, 920523”, “Slide No. 3317”, “USNM 83813”; “CRI000, 920401”, “Slide No. 3319”, “USNM 83814”; “CRI000, 920457”, “Slide No. 3291”; 3 3, “ 24 Oct.-12 Nov., 1992, D. Garcia”, “CRI000, 946628”, “Slide No. 3360”; “CRI000, 946660”, “Slide No. 3361”, “USNM 83815”; “CRI000, 946505”, “Slide No. 3310”; 8 3, “ 24 Nov.-26 Ene., 1993, J. Sihezar & G. Rodríguez”, “CRI001, 211953”, “Slide No. 3420”; “CRI001, 211945”, “Slide No. 3429”; “CRI001, 211970”, “Slide No. 3424”; “CRI001, 211901”, “Slide No. 3425”; “CRI001, 212031”, “Slide No. 3366”; “CRI001, 212052”; “Slide No. 3314”; “CRI001, 212056”, “Slide No. 3307”; “CRI001, 212048”, “Slide No. 3306”; 3 3, “ 19–27 Ene., 1993, D. Garcia”, “CRI001, 303941”, “Slide No. 3326”, “USNM 83816”; “CRI001, 303956”, “Slide No. 3342”; “CRI001, 303970”, “Slide No. 3344”; 1 3, “ 10–27 Mar., 1993, K. Taylor”, “CRI001, 319322”, “Slide No. 3345”; 2 3, “ 19–27 Ene., 1993, D. Garcia”, “CRI001, 304029”, “Slide No. 3397”; “CRI001, 303994”, “Slide No. 3393”, “USNM 83817”; 14 3, “ 21–25Mar., 1993, D. Garcia, # 2765”, “CRI001, 684196”, “Slide No. 3410”; “CRI001, 684256”, “Slide No. 3411”; “CRI001, 684202”, “Slide No. 3412”; “CRI001, 684299”, “Slide No. 3413”; “CRI001, 684211”, “Slide No. 3415”; “CRI001, 684173”, “Slide No. 3416”; “CRI001, 684343”, “Slide No. 3414”; “CRI001, 684283”, “Slide No. 3457”; “CRI001, 684278”, “Slide No. 3459”; “CRI001, 684138”, “Slide No. 3408”; “CRI001, 684139”, “Slide No. 3409”; “CRI001, 684344”, “Slide No. 3403”, “USNM 83818”; “CRI001, 684231”, “Slide No. 3401”; “CRI001, 684180”, “Slide No. 3303”; 2 3, “ 10–13 Mar., 1994, D. Garcia, # 2767”, “CRI001, 738506”, “Slide No. 3407”; “CRI001, 738470”, “Slide No. 3419”; 1 3, “ 7–19 Feb., 1994, D. Garcia, # 2624”, “CRI001, 986253”, “Slide No. 3477”, “USNM 83819”; 1 3, “ 10–18 Abr., 1994, K. Taylor, # 2838”, “CRI001, 793491”, “Slide No. 3298”; 9 3, “Est. Maritza, 600 m, lado O Volcan Orosi, Prov. Guanacaste, COSTA RICA, D. Garcia, 28 Feb.-10 Mar., 1992, L-N-326900, 373000”, “CRI000, 695284”, “Slide No. 3490”, “USNM 83820”; “CRI000, 695248”, “Slide No. 3492”; “CRI000, 695274”, “Slide No. 3493”; “CRI000, 695250”, “Slide No. 3494”; “CRI000, 695324”, “Slide No. 3495”; “CRI000, 695326”, “Slide No. 3496”; “CRI000, 695263”, “Slide No. 3510”; “CRI000, 427704”, “Slide No. 3513”; “CRI000, 695266”, “Slide No. 3838”, “USNM 83821”; 1 3, “F.A. Quesada, 27 Feb.-10 Mar. 1992 ”, “CRI000, 889095”, “Slide No. 3537”; 1 3, “R. Vargas”, “CRI000, 468682”, “Slide No. 3498”; 1 3, “I Curso Microlepidoptera, Jul. 1990 ”, “CRI000, 179547”, “Slide No. 3502”; 1 3, “II Curso Parataxonomos, Ago., 1990”, “CRI000, 680747”, “Slide No. 3522”; 1 3, “Est. Cacao, 1000–1400m, lado SO Volcan Cacao, P.N. Guanacaste, COSTA RICA, G. Rodrigues, 21–28 May., 1992, L-N-323300, 375700”, “CRI000, 684309”, “Slide No. 3279”, “Wing Slide No. 7026”; 2 3, “D. Brenes, 21–29 May., 1992”, “CRI000, 488002”, “Slide No. 3157”, “USNM 83822”; CRI000, 488042”, “Slide No. 3187”, “Wing Slide No. 7036”; 1 3, “C. Chaves, Mar. 1991 ”, “CRI000, 317627”, “Slide No. 3162”; 1 3, “C. Chaves, Abr. 1991 ”, “CRI000, 434842”, “Slide No. 3136”, “USNM 83823”; 1 3, “ June 1991 ”, “CRI000, 623350”, “Slide No. 3182”, “Wing Slide No. 7037”; 1 3, “ 21–29 May., 1992, D. Garcia”, “CRI001, 290689 ”, “Slide No. 3121”; 1 3, “ 13–17 Feb., 1995, E. Fletes, # 4766”, “CRI002, 143101”, “Slide No. 3175”, “USNM 83824”; 1 3, “II Curso Parataxonomos, Jun., 1990”, “CRI000, 661534”, “Slide No. 3137”; 1 3, “Est. Sirena, P.N. Corcovado, Prov. Puntarenas, COSTA RICA, 1–100 m, Mar., 1994, G. Fonseca, L-S- 270500, 508300, # 2769”, “CRI001, 755498”, “Slide No. 3928”; 1 3, “Mar., 1992”, “CRI000, 780351”, “Slide No. 3937”; 2 3, “G. Fonseca, Abr. 1992 ”, “CRI000, 794323”, “Slide No. 3921”; “CRI000, 794535”, “Slide No. 3923”; 2 3, “San Luis, Monteverde, Prov. Puntarenas, COSTA RICA, 1000–1350 m, Abr., 1994, Z. Fuentes, L- N-449250, 250850, # 2845”, “CRI001, 796750”, “Slide No. 3911”, “USNM 83825”; “CRI001, 796758”, “Slide No. 3906”, “USNM 83826”; 1 3, “Feb., 1995, Z. Fuentes, # 4393”, “CRI002, 165555”, “Slide No. 3859”; 1 3, “Dic., 1994, de Luz, Z. Fuentes, # 3380”, “CRI002, 119330”, “Slide No. 3857”, “USNM 83827”; “USNM 83827”; 2 3, “Tierras Morenas, Prov. Guanacaste, COSTA RICA, Mar. 1994, G. Rodriguez, L-S-283950, 424500, # 2762”, “CRI001, 683723”, “Slide No. 3582”; “CRI001, 683731”, “Slide No. 3585”; 1 3, “Fca. Jenny, 30 km N Liberia, P.N. Guanacaste, COSTA RICA, E. Araya & R. Espinoza, Oct. 1990, L-N-316200, 364400”, “CRI000, 498368”, “Slide No. 3475”; 1 3, “Quepos, 80 m P.N. Manuel Antonio, Prov. Puntarenas, COSTA RICA, R. Zuniga, Ene. 1991, L-S-370900, 448800”, “CRI000, 366807”, “Slide No. 3947”, “USNM 83828”; 1 3, “Est. La Casona, 1520 m, Res. Biol. Monteverde, Prov. Puntarenas, COSTA RICA, Dic. 1992, N. Obando, L-N-253250, 449700”, “CRI001, 358264”, “Slide No. 3738”, “USNM 83829”; 1 3, “Est. Mengo, 1100 m, SW side Volcan Cacao, Prov. Guanacaste, COSTA RICA, Feb. 1989, GNP Biodiversity Survey, 85°28'10"W, 10°55'43"N ”, “CRI001, 054909”, “Slide No. 3605”, “USNM 83830”; 13“Est. Murciélago, 8 km SO de Cuajiniquil, Prov. Guanacaste, COSTA RICA, 100 m, 7–25 Mar. 1994, F. Quesada, L-N-320300, 347200, # 2794”, “CRI001, 758757”, “Slide No. 3588” [98 in INBio, 41 in USNM].

Distribution (Map 14). Blastobasis echus is known from several collecting sites near or along the Cordillera de Guanacaste, the Cordillera de Tilarán, and the Cordillera de Talamanca; two dry-forest sites in northwestern Costa Rica, one site in the north-central part of the country near the border of Nicaragua, and two sites along eastern Pacific Coast.

Etymology. The specific epithet echus is derived from the Latin echo.

Figures

FIGURES 11 – 20. Wing venation of Blastobasinae. 11, Blastobasis erae (slide 7040). 12, Blastobasis iuanae (slide 7045). 13, Blastobasis graminea Adamski, 1999 (slide 83226). 14, Blastobasis neniae (slide 7055). 15, Blastobasis orithyia (slide 7053). 16, Blastobasis babae (slide 7049). 17, Blastobasis echus (slide 7036). 18, Blastobasis coffeaella (Busck, 1925) (USNM slide 83227). 19, Blastobasis rotullae (slide 7041). 20, Blastobasis deliciolarum (slide 7047).

FIGURES 99 – 104. Male genitalia of Blastobasinae. 99 – 100, Blastobasis echus (slide 3710). 101 – 102, Blastobasis litis (slide 3632). 103 – 104, Blastobasis chanes (slide 3450).

FIGURES 292 – 299. Male sex scales on first flagellomere of Blastobasis. 292, Blastobasis usurae. 293, Blastobasis echus. 294, Blastobasis litis. 295, Blastobasis chanes. 296, Blastobasis fax. 297, Blastobasis coffeaella (Busck, 1925). 298, Blastobasis rotullae. 299, Blastobasis custodis.

FIGURES 336 – 343. Imagos of Blastobasinae. 336, Blastobasis echus, holotype. 337, Blastobasis litis, holotype. 338, Blastobasis chanes, holotype. 339, Blastobasis fax, holotype. 340, Blastobasis coffeaella (Busck, 1925). 341, Blastobasis rotullae, holotype. 342, Blastobasis custodis, holotype. 343, Blastobasis deliciolarum, holotype.