Pinaceocladichnus onubensis Mayoral, 1988

Gaaloul, Nadia, Uchman, Alfred, Ali, Syrine Ben, Janiszewska, Katarzyna, Stolarski, Jarosław, Kołodziej, Bogusław & Riahi, Sami, 2023, In vivo and post-mortem bioerosion traces in solitary corals from the upper Pliocene deposits of Tunisia, Acta Palaeontologica Polonica 68 (4), pp. 659-681 : 671-672

Pinaceocladichnus onubensis Mayoral, 1988

Figs. 8B , 12A .

Material.—INGUJ265P151, 152, 154, 189; cylindrical, contorted tunnels in coralla; El Melah stream section, the upper part of the Argiles de Sidi Barka Formation (upper Pliocene) of Tunisia.

Diagnosis.— Pinaceocladichnus with very regular bifurcation systems. They present sub-oval or fusiform openings located very close to the corresponding tunnels that connect with elongated, cylindrical or sac-shaped chambers. They are arranged parallel to the previous tunnels (translated from Mayoral 1988).

Description. —A system of furrows, which are ca. 0.1– 0.15 mm wide, composed of straight, slightly curved, or slightly winding master furrows, and shorter straight to slightly curved furrows that branch from the same point of the master furrow (pinnate branching) under the angle 35–65°. Very small, elongated widenings are situated along or very close to the furrows.

Stratigraphic and geographic range.—Ordovician ( Estonia; Toom et al. 2019) to the Recent (worlwide).

Figures

Fig. 8. Caulostrepsis, a domichnion produced by polychates, in the skeleton of caryophylliid coral Ceratotrochus (Edwardsotrochus) duodecimcostatus Goldfuss, 1826), from the El Melah stream section, the upper part of the Argiles de Sidi Barka Formation (upper Pliocene) of Tunisia. A. Caulostrepsis cretacea (Voigt, 1971), INGUJ265P156. B. Caulostrepsis taeniola Clarke, 1908 and Pinaceocladichnus onubensis Mayoral, 1988 (Pn), INGUJ265P154. C.Caulostrepsis avipesBromleyand D’Alessandro,1983,INGUJ265P176.D.A clusterofCaulostrepsis taeniolaClarke,1908,INGUJ265P162.E–G.Caulostrepsis penicillus isp. nov. E. INGUJ265P158 holotype (a), and paratype (b). F. INGUJ265P174 paratype. G. INGUJ265P172, partial views (G1, G2).

Fig.12. Pinaceocladichnus onubensis Mayoral,1988, (a domichnion produced by ctenostome bryozoans) and associated trace fossilsin the skeleton of caryophylliid coral Ceratotrochus (Edwardsotrochus) duodecimcostatus (Goldfuss, 1826), from the El Melah stream section, the upper part of the Argiles de Sidi Barka Formation (upper Pliocene) of Tunisia.A. INGUJ265P152, different placesof the same corallum (A1,A2). B. INGUJ265P154,also Maeandropolydora elegans Bromley and D’Alessandro, 1983 (Me).C. INGUJ265P151,C1, C2 different places of the same corallum, also Caulostrepsis cretacea (Voigt,1971) (Cc).