Komiyandra niisatoi, Santos-Silva & Heffern & Matsuda, 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 31

Komiyandra niisatoi sp. nov.

( Fig. 62, 63 , 111 , 184, 185 , 251 , 294 , 338 , 460-462 )

Etymology. Dedicated to our colleague, Dr. Tatsuya Niisato, of Japan, who has extensively published on Cerambycidae , and provided important specimens for this study.

Type material. Holotype M, INDONESIA, Sulawesi, West Sulawesi: between Tabone and Mamasa, II.2001, ( MZSP – donated by Ziro Komiya) . Paratype (2 M, 2 F), as follows: INDONESIA, Sulawesi, North Sulawesi: Tondano, M, III.1985, (no collector indicated) ( ZKCO) ; 1 M, 1 F, III.1985, (no collector indicated) ( KMCT) ; Utara, Dumoga-Bone National Park (Sub-camp Edwards; 1140m), F, X.1985, J. Van Stalle coll. ( IRSN) .

Description. Integument very dark-brown; head and mandibles almost fully black.

Male ( Fig. 460 ). Dorsal face of head, on gibbosities, moderately, coarsely and abundantly punctate; area between gibbosities and occiput with punctures similar to that on gibbosities, but with an impunctate area at each side of central region; area between gibbosities and ocular carina clearly depressed, almost fully smooth; ocular carina elevated, clearly bifurcated in “Y” near posterior edge of eyes ( Fig. 460 ). Eyes ( Fig. 111 ) moderately narrow; posterior ocular edge ( Fig. 460 ) very distinct. Central area of clypeus almost vertical, with lateral depressions. Central projection of labrum ( Fig. 62 ) wide and truncate at apex. Submentum depressed; somewhat coarsely, sparsely punctate; pilosity short and very sparse; anterior edge elevated throughout. Mandibles approximately as long as head; teeth of inner margin ( Fig. 184 ) placed approximately at middle. Ventral sensorial area of antennomeres III-XI not visible from side ( Fig. 251 ), and not divided by carina.

Pronotum finely, sparsely punctate at central region, clearly coarser and more abundant laterally, mainly near anterior angles; anterior edge slightly sinuous; anterior angles not projected forward; lateral angles marked and widely obtuse; posterior angles obtuse. Basal two-thirds of elytra with punctures barely coarse near suture, and coarser and more abundant laterally; apical third with punctures just barely coarse and abundant; each elytron with two carinae. Metasternum punctation coarse and moderately sparse laterally, finer and sparser towards the metasternal suture. Metafemur ( Fig. 461 ) moderately narrow and elongated. Dorsal face of metatibia flat. Metatarsomere V (without claws) longer than I-III together ( Fig. 294 ).

Female ( Fig. 462 ). Central region of the clypeus, close to front, strongly oblique, without depressions at sides. Central area of head, between the gibbosities and occiput fully punctate. Central projection of labrum as in Fig. 63 . Mandibles as in Fig. 185 .

Variability. Male: central projection of labrum wide and truncate at apex. Female: head and mandibles dark-brown with blackish areas.

Dimensions in mm (M / F). Total length (including mandibles), 22.5/21.0; prothorax: length, 4.6/4.1; anterior width, 5.7/4.8; posterior width, 4.4/4.6; humeral width, 5.7/5.9; elytral length, 12.5/13.0.

Comments. Komiyandra niisatoi is similar to K. javana ( Fig. 419, 421 ). It differs by the: integument darker; ocular carina clearly bifurcated near the posterior edge of the eyes; central area of the clypeus of the male, close to the front, with depressions at the sides; central area of the clypeus of females strongly oblique. In K. javana , the integument is lighter brown, the ocular carina is not bifurcated or the bifurcation is barely indicated, the clypeus of males, close to the front, has no depressions to the sides, and in females, the central area of the clypeus is less oblique. It differs from K. philippinensis , mainly, by the darker integument and slimmer body.

Figures

Figure 45-74. 45-73) Labrum. 45) Komiyandra luzonica, holotype male. 46) K. luzonica paratype female. 47) K. philippinensis, paratype male. 48) K. philippinensis, paratype female. 49) K. mindanao, holotype male. 50) K. mindanao, paratype female. 51) K. mehli, holotype male. 52) K. mehli, paratype female. 53) K. lombokia, holotype male. 54) K. lombokia, paratype female. 55) K. sulawesiana, holotype male. 56) K. sulawesiana, paratype female. 57) K. irianjayana, holotype male. 58) K. menieri, paratype male. 59) K. menieri, paralectotype female of P. janus. 60) K. sangihe, holotype male. 61) K. mindoro, holotype male. 62) K. niisatoi, holotype male. 63) K. niisatoi, paratype female. 64) K. drumonti, holotype male. 65) K. cabigasi, paratype male. 66) K. cabigasi, paratype female. 67) K. koni, paratype male. 68) K. koni, paratype female. 69) K. johkii, holotype male. 70) K. poggii, holotype male. 71) K. poggii, paratype female. 72) K. uenoi, paratype male. 73) K. uenoi, paratype female. 74) Hawaiiandra puncticeps, head dorsal view, female (CL, clypeus; LB, labrum; FR, frons; at, anterior tentorial pit; cl, clypeolabral suture; es, epistomal suture).

Figure 105-117. Head, lateral view. 105) K. lombokia, paratype male. 106) K. sulawesiana, holotype male. 107) K. irianjayana, holotype male. 108) K. menieri, paratype male. 109) K. sangihe, paratype male. 110) K. mindoro, holotype male. 111) K. niisatoi, paratype male. 112) K. drumonti, holotype male. 113) K. cabigasi, paratype male. 114) K. koni, paratype male. 115) K. johkii, holotype male. 116) K. poggii, holotype male. 117) K. uenoi, paratype male.

Figure 177-199. 177-195) Mandibles, dorsal view. 177) Komiyandra sulawesiana, holotype male. 178) K. sulawesiana, paratype female. 179) K. irianjayana, holotype male. 180) K. menieri, paratype male. 181) K. menieri, paratype female (paralectotype of Parandra janus). 182) K. sangihe, paratype male. 183) K. mindoro, holotype male. 184) K. niisatoi, holotype male. 185) K. niisatoi, paratype female. 186) K. drumonti, holotype male. 187) K. cabigasi, paratype male. 188) K. cabigasi, paratype female. 189) K. koni, paratype male. 190) K. koni, paratype female. 191) K. johkii, holotype male. 192) K. poggii, holotype male. 193) K. poggii, paratype female. 194) K. uenoi, paratype male. 195) K. uenoi, paratype female. 196-199) Maxillary palp. 196) Komiyandra javana, paratype male. 197) Caledonandra austrocaledonica, male. 198) C. passandroides, male. 199) Hawaiiandra puncticeps, male.

Figure 235-251. Antenna. 235) Komiyandra formosana, male. 236) K. lanyuana, male. 237) K. javana, holotype male. 238) K. nayani, paratype male. 239) K. ohbayashii, holotype male. 240) K. luzonica, paratype male. 241) K. philippinensis, paratype male. 242) K. mindanao, holotype male. 243) K. mehli, paratype male. 244) K. vivesi, holotype male. 245) K. lombokia, paratype male. 246) K. sulawesiana, paratype male. 247) K. irianjayana, holotype male. 248) K. menieri, paratype male. 249) K. sangihe, holotype male. 250) K. mindoro, holotype male. 251) K. niisatoi, holotype male.

Figure 277-299. Metatarsus. 277) Komiyandra shibatai, male. 278) K. formosana, male. 279) K. lanyuana, male. 280) K. javana, paratype male. 281) K. nayani, paratype male. 282) K. ohbayashii, holotype male. 283) K. luzonica, paratype male. 284) K. philippinensis, paratype male. 285) K. mindanao, holotype male. 286) K. mehli, paratype male. 287) K. vivesi, holotype male. 288) K. lombokia, paratype male. 289) K. sulawesiana, holotype male. 290) K. irianjayana, holotype male. 291) K. menieri, holotype male. 292) K. sangihe, holotype male. 293) K. mindoro, holotype male. 294) K. niisatoi, holotype male. 295) K. drumonti, holotype male. 296) K. cabigasi, paratype male. 297) K. koni, paratype male. 298) K. poggii, holotype male. 299) K. uenoi, paratype male.

Figure 335-340. Maps. 335) Komiyandra mehli. 336) K. sulawesiana. 337) K. vivesi (circle), K. mindanao (square). 338) K. niisatoi (circle), K. janus (square), K. menieri (triangle). 339) K. sangihe. 340) K. lombokia.

Figure 454-460. Habitus. 454) Komiyandra irianjayana, holotype male, punctation, head and pronotum. 455) K. menieri, paratype female, dorsal view (paralectotype of Parandra janus). 456) K. sangihe, holotype male, dorsal view. 457) K. sangihe, holotype male, ventral view. 458) K. mindoro, holotype male, dorsal view. 459) K. mindoro, holotype male, ventral view. 460) K. niisatoi, holotype male, dorsal view.

Figure 461-466. Habitus. 461) Komiyandra niisatoi, holotype male, ventral view. 462) K. niisatoi, paratype female, dorsal view. 463) K. drumonti, holotype male, dorsal view. 464) K. drumonti, holotype male, ventral view. 465) K. drumonti, holotype male, lateral view. 466) K. cabigasi, holotype male, dorsal view.

Figure 416-421. Habitus. 416) Komiyandra lanyuana, male, dorsal view. 417) K. lanyuana, male, ventral view. 418) K. lanyuana, female, dorsal view. 419) K. javana, holotype male, dorsal view. 420) K. javana, holotype male, ventral view. 421) K. javana, paratype female, dorsal view.