Komiyandra, Santos-Silva & Heffern & Matsuda, 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 7-8

Komiyandra , new genus

Etymology. Dedicated to our colleague, Mr. Ziro Komiya (Komiya + Parandra ), of Japan, who has published numerous papers on Oriental Prioninae and provided many of the important specimens for this study. Feminine gender.

Type species. Parandra janus Bates, 1875 .

Description. Dorsal area of head, between eyes, with gibbosities well marked, separated by furrow deep or barely deep, without central depression in “V”. Ocular carina elevated and distinct from middle to clypeus. Eyes ( Fig. 94 ) narrow (maximum width equal to 0.4 times the total length); posterior ocular edge ( Fig. 452 ) prominent or very prominent; anterior ocular edge with concavity well marked. Frontoclypeal suture just visible laterally (sometimes absent). Central region of clypeus vertical, oblique or strongly oblique. Clypeolabral suture visible in full extension or only laterally. Central projection of male labrum varying from wide and truncate at apex to narrow and sharpened apically; central projection of female usually as in male, but when truncate, always narrower. Mandibles of major males ( Fig. 177, 184 ) subfalciform (almost identical to female in minor males), shorter than head or slightly shorter, wide at base of latero-outer face ( Fig. 114 ); dorsal carina elevated, well marked from base to apical third; inner margin with two teeth, together protracted, located in middle or after middle; apex with two large teeth, visible dorsally, and a third, small, not visible dorsally; outer face without large tooth around middle ( Fig. 107 ). Mandibles of females ( Fig. 169 , 185 ) Birandra -like, wide at base of latero-outer face; dorsal carina elevated only at basal third, ending approximately in middle; inner margin with two teeth together protracted, located in middle; apex and outer face as in males. Mentum with hair long and sparse. Galea ( Fig. 196 , 200 ) long (surpassing the apex of second segment of maxillary palp). Ventral sensorial area of antennae ( Fig. 235, 245 ) not visible from side, divided ( Fig. 312, 313 ) or not by low carina, not visible from side (sometimes visible from the side on antennomere XI or X-XI); ventral sensorial area of antennomere XI not extending into dorsal area; dorsal sensorial area of antennomere XI small, well delimited, not divided by carina.

Pronotum strongly convex in apical half (close to head); anterior edge barely sinuous (mainly in males) to sinuous (mainly in females); anterior angles projected to front (usually more distinct in females); lateral angle very distinct and obtuse, or slightly distinct and obtuse (sometimes variable intraspecific); posterior angles distinct and obtuse (sometimes almost in right angle). Elytra punctate, usually distinctly and abundantly. Veins MP 3 and MP 4 fused at their apex ( Fig. 211 ). Apex of prosternal process barely enlarged. Femora glabrous or sub-glabrous. Dorsal face of tibiae rounded, or flat or shallowly furrowed. Procoxal cavities clearly open behind. Paronychium with one seta (two setae in K. ohbayashii sp. nov.).

Included species. Komiyandra janus ( Bates, 1875) , comb. nov.; K. shibatai ( Hayashi, 1963) , comb. nov.; K. formosana ( Miwa and Mitono, 1939) , comb. nov.; K. lanyuana ( Hayashi, 1981) , comb. nov.; K. javana Santos-Silva, Heffern and Matsuda , sp. nov.; K. nayani Santos-Silva, Heffern and Matsuda , sp. nov.; K. ohbayashii Santos-Silva, Heffern and Matsuda , sp. nov.; K. luzonica Santos-Silva, Heffern and Matsuda , sp. nov.; K. philippinensis Santos-Silva, Heffern and Matsuda , sp. nov.; K. mindanao Santos-Silva, Heffern and Matsuda , sp. nov.; K. mehli Santos-Silva, Heffern and Matsuda , sp. nov.; K. vivesi Santos-Silva, Heffern and Matsuda , sp. nov.; K. lombokia Santos-Silva, Heffern and Matsuda , sp. nov.; K. sulawesiana Santos-Silva, Heffern and Matsuda , sp. nov.; K. irianjayana Santos-Silva, Heffern and Matsuda , sp. nov.; K. menieri Santos-Silva, Heffern and Matsuda , sp. nov.; K. sangihe Santos-Silva, Heffern and Matsuda, sp. nov.; K. mindoro Santos-Silva, Heffern and Matsuda , sp. nov.; K. niisatoi Santos-Silva, Heffern and Matsuda , sp. nov.; K. drumonti Santos-Silva, Heffern and Matsuda , sp. nov.; K. cabigasi Santos-Silva, Heffern and Matsuda , sp. nov.; K. koni Santos-Silva, Heffern and Matsuda , sp. nov.; K. johkii Santos-Silva, Heffern and Matsuda , sp. nov.; K. poggii Santos-Silva, Heffern and Matsuda , sp. nov.; and K. uenoi Santos-Silva, Heffern and Matsuda , sp. nov.

Geographical distribution ( Fig. 316 ). Japan (Ryukyu Islands), Taiwan, Philippines, Malaysia (Borneo: Sabah), Indonesia (Borneo, Sulawesi, Ternate, Java, Lombok, Sumbawa, Moluccas, Irian Jaya, Sumatra, Sangihe, Seram).

Comments. Komiyandra differs from Birandra Santos-Silva, 2002 , and Acutandra Santos-Silva, 2002 , by mandibles of major males being sub-falciform ( Fig. 162 ), and by veins MP 3 and MP 4 ( Fig. 211 ) fused together at apices. In Birandra , mandibles of major males ( Fig. 118 ) are clearly falciform and veins MP 3 and MP 4 ( Fig. 212 ) are separate at apices. In Acutandra , mandibles of major males ( Fig. 120 ) are very similar to that of females (not falciform or sub-falciform), and veins MP 3 and MP 4 ( Fig. 205 ) are as in Birandra . Different from Archandra Lameere, 1912 , and Parandra Latreille, 1802 , because these genera have distinctly falciform mandibles in major males and veins MP 3 and MP 4 ( Fig. 207, 208 ) not fused together at apices, and by procoxal cavities closed behind. From Neandra Lameere, 1912 , different by procoxal cavities open behind, and presence of paronychium clearly exposed. In Neandra , procoxal cavities are closed behind, and paronychium not exposed. Different from Stenandra Lameere, 1912 , by procoxal cavities open behind, and by the paronychium clearly exposed (closed and not exposed in Stenandra ). See comments on Melanesiandra , Hawaiiandra , Caledonandra , Malukandra , Papuandra , and Storeyandra .

Figures

Figure 90-104. Head, lateral view. 90) Papuandra queenslandensis, holotype female. 91) P. rothschildi, holotype male. 92) P. oberthueri, holotype female. 93) Stenandra kolbei, female. 94) Komiyandra shibatai, male. 95) K. formosana, male. 96) K. lanyuana, male. 97) K. javana, holotype male. 98) K. nayani, paratype male. 99) K. ohbayashii, holotype male. 100) K. luzonica, paratype male. 101) K. philippinensis, paratype male. 102) K. mindanao, holotype male. 103) K. mehli, paratype male. 104) K. vivesi, holotype male.

Figure 447-453. Habitus. 447) Komiyandra sulawesiana, paratype female, dorsal view. 448) K. irianjayana, holotype male, dorsal view. 449) K. irianjayana, holotype male, ventral view. 450) K. menieri, paratype male, dorsal view. 451) K. menieri, paratype male, ventral view. 452) K. menieri, paratype male, punctation, head and pronotum. 453) K. philippinensis, paratype male, punctation, head and pronotum.

Figure 177-199. 177-195) Mandibles, dorsal view. 177) Komiyandra sulawesiana, holotype male. 178) K. sulawesiana, paratype female. 179) K. irianjayana, holotype male. 180) K. menieri, paratype male. 181) K. menieri, paratype female (paralectotype of Parandra janus). 182) K. sangihe, paratype male. 183) K. mindoro, holotype male. 184) K. niisatoi, holotype male. 185) K. niisatoi, paratype female. 186) K. drumonti, holotype male. 187) K. cabigasi, paratype male. 188) K. cabigasi, paratype female. 189) K. koni, paratype male. 190) K. koni, paratype female. 191) K. johkii, holotype male. 192) K. poggii, holotype male. 193) K. poggii, paratype female. 194) K. uenoi, paratype male. 195) K. uenoi, paratype female. 196-199) Maxillary palp. 196) Komiyandra javana, paratype male. 197) Caledonandra austrocaledonica, male. 198) C. passandroides, male. 199) Hawaiiandra puncticeps, male.

Figure 105-117. Head, lateral view. 105) K. lombokia, paratype male. 106) K. sulawesiana, holotype male. 107) K. irianjayana, holotype male. 108) K. menieri, paratype male. 109) K. sangihe, paratype male. 110) K. mindoro, holotype male. 111) K. niisatoi, paratype male. 112) K. drumonti, holotype male. 113) K. cabigasi, paratype male. 114) K. koni, paratype male. 115) K. johkii, holotype male. 116) K. poggii, holotype male. 117) K. uenoi, paratype male.

Figure 148-176. Mandibles, dorsal view. 148) Papuandra norfolkensis, holotype male (left mandible). 149) P. norfolkensis, holotype male (right mandible). 150) P. norfolkensis, paratype female. 151) P. rothschildi, holotype male. 152) P. rothschildi, paratype female. 153) Komiyandra janus, female. 154) K. shibatai, male. 155) K. shibatai, female. 156) K. formosana, male. 157) K. formosana, female. 158) K. lanyuana, male. 159) K. lanyuana female. 160) K. javana, holotype male. 161) K. javana, paratype female. 162) K. nayani, paratype male. 163) K. nayani, paratype female.164) K. ohbayashii, holotype male. 165) K. ohbayashii, paratype female. 166) K. luzonica, holotype male. 167) K. luzonica, paratype female. 168) K. philippinensis, holotype male. 169) K. philippinensis, paratype female. 170) K. mindanao, holotype male. 171) K. mindanao, paratype female. 172) K. mehli, holotype male. 173) K. mehli, paratype female. 174) K. vivesi, holotype male. 175) K. lombokia, holotype male. 176) K. lombokia, paratype female.

Figure 200-209. 200-204) Maxillary palp. 200) Komiyandra nayani, paratype male. 201) Papuandra araucariae, male. 202) Storeyandra frenchi, female. 203) Melanesiandra striatifrons, male. 204) Acutandra murrayi, female. 205-209) Wing. 205) Acutandra murrayi, female. 206) Neandra brunnea, male. 207) Archandra caspia, male. 208) Parandra (Parandra) ubirajarai, male. 209) Hawaiiandra puncticeps, male.

Figure 235-251. Antenna. 235) Komiyandra formosana, male. 236) K. lanyuana, male. 237) K. javana, holotype male. 238) K. nayani, paratype male. 239) K. ohbayashii, holotype male. 240) K. luzonica, paratype male. 241) K. philippinensis, paratype male. 242) K. mindanao, holotype male. 243) K. mehli, paratype male. 244) K. vivesi, holotype male. 245) K. lombokia, paratype male. 246) K. sulawesiana, paratype male. 247) K. irianjayana, holotype male. 248) K. menieri, paratype male. 249) K. sangihe, holotype male. 250) K. mindoro, holotype male. 251) K. niisatoi, holotype male.

Figure 300-314. 300-303) Metatarsomere V. 300) Komiyandra shibatai, male. 301) K. formosana, male. 302) K. lanyuana, male. 303) K. uenoi, holotype male. 304-311) Aedeagus. 304) K. shibatai, parameres. 305) K. shibatai, median lobe. 306) K. formosana, parameres. 307) K. formosana, median lobe. 308) K. lanyuana, parameres. 309) K. lanyuana, median lobe. 310) K. uenoi, paratype parameres. 311) K. uenoi, paratype median lobe. 312-314) Antennomere XI. 312) K. nayani, paratype male. 313) K. javana, holotype male. 314) Melanesiandra striatifrons, male.

Figure 210-217. 210-214) Wings. 210) Papuandra araucariae, male. 211) Komiyandra nayani, paratype male. 212) Birandra (Birandra) silvaini, male. 213) Caledonandra passandroides, male. 214) Melanesiandra striatifrons, male. 215-217) Antenna. 215) Stenandra kolbei, female. 216) Hawaiiandra puncticeps, male. 217) Storeyandra frenchi, male.

Figure 315-316. Maps. 315) Distribution of Melanesiandra, Caledonandra, Storeyandra, Hawaiiandra, Malukandra, and Stenandra in Asia. 316) Distribution of Komiyandra and Papuandra.

Figure 118-147. Mandibles, dorsal view. 118) Birandra (Yvesandra) angulicollis, male. 119) B. (Y.) angulicollis, female. 120) Acutandra araucana, male. 121) Parandra (Parandra) glabra, female. 122) Stenandra kolbei, female. 123) Hawaiiandra puncticeps, male. 124) Storeyandra frenchi, male. 125) S. frenchi, female. 126) Malukandra heterostyla, holotype male. 127) M. jayawijayana, holotype male. 128) M. hornabrooki, holotype male. 129) M. hornabrooki, paratype female. 130) Caledonandra austrocaledonica, male. 131) C. austrocaledonica, female. 132) C. passandroides, male. 133) C. passandroides, female. 134) Melanesiandra striatifrons, male. 135) M. striatifrons, female. 136) M. solomonensis, male. 137) M. solomonensis, female. 138) M. bougainvillensis, holotype male. 139) M. bougainvillensis, paratype female. 140) M. birai, holotype male. 141) Papuandra araucariae, paratype male. 142) P. araucariae, female. 143) P. gressitti, holotype male. 144) P. weigeli, holotype male. 145) P. weigeli, paratype female. 146) P. queenslandensis, holotype male. 147) P. oberthueri, holotype female.