Komiyandra janus ( Bates, 1875 ), 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 21-23

Komiyandra janus ( Bates, 1875)

( Fig. 31 , 153 , 233 , 276 , 338 , 405-409 )

Parandra janus Bates, 1875: 47 .

Birandra (Birandra) janus ; Santos-Silva and Shute 2009: 32.

Description. Integument dark-brown; parts of head and of mandibles blackish; legs pale-brown with apex of femurs dark-brown.

Male ( Fig. 405 ). Dorsal face of head, on gibbosities, coarsely and abundantly punctate; area between gibbosities and occiput less coarsely and abundantly punctate; gibbosities well marked, separated by wide furrow moderately deep; area between gibbosities and ocular carina depressed, almost fully smooth; ocular carina elevated, slightly bifurcated in “Y” near posterior edge of eyes ( Fig. 407 ); area behind eyes just coarsely, not confluently punctate, except region close to eyes that is smooth. Eyes moderately wide; posterior ocular edge prominent ( Fig. 407 ), without abrupt declivity. Central area of clypeus strongly oblique close to front. Labrum narrow, not projected laterally; central projection of labrum ( Fig. 407 ) narrow, subacute at apex. Submentum depressed, punctation coarse and sparse, more abundant close to clypeus; pilosity very short and very sparse; anterior edge elevated just at lateral. Teeth of inner margin of mandibles ( Fig. 407 ) placed at apical half. Ventral sensorial area of antennomeres III-XI not visible from side, divided by low carina, slightly visible from side in antennomeres X-XI.

Pronotum finely punctate at central region, gradually coarser and more abundant towards anterior and lateral margins, mainly near anterior angles; anterior edge ( Fig. 407 ) slightly sinuous; anterior angles rounded, not projected forward; lateral angles absent; posterior angles distinct, obtuse. Elytra coarsely and abundantly punctate, gradually finer and more abundant at apical third. Metasternum with coarse and sparse punctation, finer and sparser towards metasternal suture. Metafemur ( Fig. 406 ) short and enlarged. omeres III-XI not visible from the side, divided by low carina, slightly visible from the si

Female ( Fig. 409 ). Dorsal face of head, on gibbosities, sparsely punctate; gibbosities separated by wide, shallow furrow, with punctiform depression approximately at middle. Central projection of labrum ( Fig. 31 ) as in male. Mandible ( Fig. 153 ). Antenna ( Fig. 233 ). Pronotum finer and sparser than in male. Dorsal face of metatibia rounded. Metatarsomere V (without claws) longer than I-III together ( Fig. 276 ).

Dimensions in mm (F). Total length (including mandibles), 17.0/18.8; prothorax: length, 3.5/3.5; anterior width, 4.5/4.0; posterior width, 3.5/4.0; humeral width, 4.4/4.9; elytral length, 10.3/11.3.

Dimensions in lines (syntypes M / F) according to Bates (1875): 11 (approximately 23.3 mm).

Geographical distribution ( Fig. 338 ). Indonesia (Sulawesi). Komiyandra janus has been recorded from Japan, Taiwan, Indonesia (Java, Sulawesi, Moluccas, Ternate, New Guinea - Irian Jaya), and Philippines (Luzon, Mindanao). However, we believe that this species only occurs in Sulawesi. Thus, all previous distributional records for K. janus should be disregarded, except Sulawesi.

Material examined. INDONESIA, Sulawesi, North Sulawesi, Utara, Gunung Mooat (1050m; station 062), F, J. Van Stalle coll. ( IRSN) ; Lake Mooat (1100 m; 20 km E Kotamobagu, 124,5 oE, 1 oN, east of Dumoga Bone National Park ), M, IX.12.1998, K. Maekawa coll. ( KMTC) .

Types, type localities. Bates (1875) described Parandra janus based on two syntypes: “One example ([male symbol]) from Dr. Meyer’s collection, Menado [sic], Celebes; one [female symbol], Andai, New Guinea (Signor D’Albertis)”. Apparently, Bates retained the male syntype for his collection and that specimen is in MNHN. According to Cambefort (2007), Oberthür succeeded in acquiring most of the Bates Collection after his death except Endomychidae , Elateridae and perhaps those from Biologia Centrali-Americana. There is no doubt that the male specimen deposited in MNHN is the true syntype. The calligraphy of one of the handwritten labels, fixed on the specimen, agrees perfectly with that of Bates, figured in Horn and Kahle (1935: XI, 11). According to Gérard L. Tavakilian (pers. comm.) the female syntype was not found in the Collection of MNHN, and according to Horn and Kahle (1935), the D’Albertis Collection is deposited at MCGD. Despite the statement of Lameere (1902), the syntype female never was in Oberthür’s Collection, and actually belongs to MCGD.

As alluded to in the “Geographical distribution”, we believe that the female syntype is not the same species as the male syntype. In order to guarantee the stability and identification of the species, we here designate as LECTOTYPE the male syntype, deposited at MNHN that has the following labels ( Fig. 408 ): 1. White: Parandra janus Bates (handwritten); (handwritten male symbol); Det. K. Matsuda, 1996 (printed, except “96”);

2. Red: Holotype (printed);

3. Gray: Type (printed);

4. White: Museum Paris Coll. H. W. Bates 1952 (printed);

5. White: Janus Bates (handwritten);

6. White: Menado Celebes (handwritten);

7. White: Ex-Musaeo H. W. Bates 1892 (printed);

8. Red: Lectotype (printed) – at present added by us.

The Paralectotype female, deposited at MCGD, has the following labels:

1. White: Andai / Ag. 1872 (handwritten); D’Albertis (printed);

2. Salmon: SYNTYPUS (printed); Parandra janus / [female symbol] / Bates, 1875 (handwritten);

3. White: Parandra janus Bates / teste Lameere 1902 (printed);

4. White: Museo Civico di Genova (printed);

5. Red: Paralectotype (printed) (added by us);

6. Red: Paratype (printed) (added by us);

7. White: Komiyandra menieri (added by us).

Comments. All bibliographic citations for Parandra janus after Bates (1875) refer to different species that, at most, mentioned the localities recorded in the original description. So, it becomes unviable to mention them in the bibliographical list.

For this study, we received many specimens incorrectly identified as Parandra janus , and unfortunately, among all these specimens only a couple related below correspond to the true K. janus . The examination of specimens identified by authors like Lameere (1902), Arigony (1984), and Hüdepohl (1990), shows that these authors confused the species and identified more than one species under the epithet “ janus ”, like Bates (1875). See comments on K. javana and K. sulawesiana .

It is impossible to know which species Gressitt (1951, 1959) confused with K. janus . His descriptions in the keys in both works do not lead to the correct identification. However, the locality recorded by Gressitt (1951) for P. janus [“Botel-Tobago I. 5 (Kotosho), near Formosa ”], based in Kano (1938), corresponds to that of K. lanyuana .

In general appearance, K. janus is similar to K. nayani , from which it differs by the sparser elytral punctation, and by the form of the area behind the eyes, that is very similar to that of Papuandra araucariae , this is, the smooth part close to the posterior ocular edge being large and not abruptly sloping. In K. nayani , the elytral punctation is very abundant, and the smooth area behind the eyes is smaller and abruptly sloping. Besides, the ventral sensorial area of antennomeres I-III is divided by distinct carina in K. janus . In K. nayani , the ventral sensorial area of the antennomeres is not divided by carina, or the carina is vaguely evident.

Figures

Figure 1-44. Labrum. 1) Papuandra araucariae, male. 2) P. araucariae, female. 3) P. gressitti, holotype male. 4) P. weigeli, holotype male. 5) P. weigeli, paratype female. 6) P. queenslandensis, holotype female. 7) P. norfolkensis, holotype male. 8) P. norfolkensis, paratype female. 9) P. rothschildi, holotype male. 10) P. rothschildi, paratype female. 11) P. oberthueri, holotype female. 12) Caledonandra austrocaledonica, male. 13) C. austrocaledonica, female. 14) C. passandroides, male. 15) C. passandroides, female. 16) Storeyandra frenchi, male. 17) S. frenchi, female. 18) Malukandra heterostyla, holotype male. 19) M. jayawijayana, holotype male. 20) M. hornabrooki, holotype male. 21) M. hornabrooki, paratype female. 22) Melanesiandra striatifrons, male. 23) M. striatifrons, holotype female of Parandra vitiensis. 24) M. solomonensis, male. 25) M. solomonensis, female. 26) M. bougainvillensis, holotype male. 27) M.bougainvillensis, paratype female.28) M.birai, holotype male. 29) Stenandra kolbei, female. 30) Hawaiiandra puncticeps, male. 31) Komiyandra janus, female. 32) K. shibatai, male. 33) K. shibatai, female. 34) K. formosana, male. 35) K. formosana, female. 36) K. lanyuana, male. 37) K. lanyuana, female. 38) K. javana, holotype male. 39) K. javana, paratype female. 40) K. nayani, paratype male. 41) K. nayani, paratype female. 42) K. ohbayashii, holotype male. 43) K. ohbayashii, paratype female. 44) K. vivesi, holotype male.

Figure 148-176. Mandibles, dorsal view. 148) Papuandra norfolkensis, holotype male (left mandible). 149) P. norfolkensis, holotype male (right mandible). 150) P. norfolkensis, paratype female. 151) P. rothschildi, holotype male. 152) P. rothschildi, paratype female. 153) Komiyandra janus, female. 154) K. shibatai, male. 155) K. shibatai, female. 156) K. formosana, male. 157) K. formosana, female. 158) K. lanyuana, male. 159) K. lanyuana female. 160) K. javana, holotype male. 161) K. javana, paratype female. 162) K. nayani, paratype male. 163) K. nayani, paratype female.164) K. ohbayashii, holotype male. 165) K. ohbayashii, paratype female. 166) K. luzonica, holotype male. 167) K. luzonica, paratype female. 168) K. philippinensis, holotype male. 169) K. philippinensis, paratype female. 170) K. mindanao, holotype male. 171) K. mindanao, paratype female. 172) K. mehli, holotype male. 173) K. mehli, paratype female. 174) K. vivesi, holotype male. 175) K. lombokia, holotype male. 176) K. lombokia, paratype female.

Figure 218-234. Antenna. 218) Malukandra jayawijayana, holotype male. 219) M. hornabrooki, paratype female. 220) Caledonandra austrocaledonica, male. 221) C. passandroides, male. 222) Melanesiandra striatifrons, male. 223) M. solomonensis, male. 224) M. bougainvillensis, holotype male. 225) M. birai, holotype male.226) Papuandra araucariae, male. 227) P. gressitti, holotype male. 228) P. weigeli, holotype male. 229) P. queenslandensis, holotype female. 230) P. norfolkensis, holotype male. 231) P. rothschildi, paratype male. 232) P. oberthueri, holotype female. 233) Komiyandra janus, female. 234) K. shibatai, male.

Figure 252-276. 252-257) Antenna. 252) Komiyandra drumonti, holotype male. 253) K. cabigasi, paratype male. 254) K. johkii, holotype male. 255) K. poggii, holotype male. 256) K. koni, paratype male. 257) K. uenoi, paratype male. 258-276) Metatarsus. 258) Stenandra kolbei, female. 259) Hawaiiandra puncticeps, male. 260) Storeyandra frenchi, male. 261) Malukandra jayawijayana, holotype male. 262) M. hornabrooki, holotype male. 263) Caledonandra austrocaledonica, male. 264) C. passandroides, male. 265) Melanesiandra striatifrons, male. 266) M. solomonensis, male. 267) M. bougainvillensis, holotype male. 268) M. birai, holotype male. 269) Papuandra araucariae, male. 270) P. gressitti, holotype male. 271) P. weigeli, holotype male. 272) P. queenslandensis, holotype female. 273) P. norfolkensis, holotype male. 274) P. rothschildi, holotype male. 275) oberthueri, holotype female. 276) Komiyandra janus, female.

Figure 335-340. Maps. 335) Komiyandra mehli. 336) K. sulawesiana. 337) K. vivesi (circle), K. mindanao (square). 338) K. niisatoi (circle), K. janus (square), K. menieri (triangle). 339) K. sangihe. 340) K. lombokia.

Figure 403-409. Habitus. 403) Papuandra oberthueri, holotype female, dorsal view. 404) P. oberthueri, holotype female, ventral view. 405) Komiyandra janus, lectotype male, dorsal view. 406) K. janus, lectotype male, ventral view. 407) K. janus, lectotype male, head and prothorax. 408) K. janus, lectotype male, labels. 409) K. janus, female, dorsal view.