Papuandra, Santos-Silva & Heffern & Matsuda, 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 63-64

Papuandra , new genus

Etymology. Papua + Parandra , in reference to Papua New Guinea, a country occupying the eastern half of the island of New Guinea. Feminine gender.

Type species. Parandra araucariae Gressitt, 1959 .

Description. Dorsal area of head, between eyes, in both sexes, with gibbosities well defined, separated by furrow deep or moderately shallow (sometimes variable intraspecifically), without central depression in “V”. Ocular carina elevated, narrow or wide, evident from middle to clypeus; in male, slightly bifurcated in “Y” near posterior edge of eyes or not bifurcated, and not bifurcated in female. Eyes narrow ( Fig. 87 ) or moderately narrow ( Fig. 86 ) in male, and somewhat or slightly wider in female; posterior ocular edge of male distinct ( Fig. 390 ) or barely so ( Fig. 387 ), and barely distinct in female; anterior ocular edge emarginate. Frontoclypeal suture not distinct or visible only laterally. Central region of clypeus oblique, strongly oblique, or vertical. Clypeolabral suture visible laterally or indicated by darkening of integument in area of suture. Central projection of male labrum wide or narrow, rounded or subtruncate at apex; in female narrow and rounded at apex. Mandibles of male sub-falciform ( Fig. 143 , 392 ) or distinctly not falciform ( Fig. 141 ), approximately same length as head, wide at base of outer margin; dorsal carina wide and not notably elevated, or narrow and clearly elevated; inner margin with two teeth together protracted (completely fused or sub-fused at apex), placed at apical third, or two teeth together protracted and clearly separated at apex, placed near middle; apex with two large teeth, visible dorsally, and a third, small tooth, not visible dorsally; outer margin ( Fig. 86 ) without tooth around middle. Mandibles of female ( Fig. 142 ) Birandra -like, wide at base of outer margin; dorsal carina wide and elevated only at basal third; inner margin with two teeth together protracted, placed at middle; apex and outer margin as in male. Mentum with hair moderately long, sparse or very sparse. Galea long ( Fig. 201 ) (surpassing the apex of second segment of maxillary palp). Ventral sensorial area of antennae ( Fig. 226 ) visible or not from side, divided or not by carina; ventral sensorial area of antennomere XI not extending to dorsal area; dorsal sensorial area of antennomere XI small or moderately large, well defined, not divided by carina, or absent.

Pronotum clearly convex, mainly on anterior half; anterior edge of male slightly concave or slightly sinuous on central region, and sinuous in female; anterior angles of male distinctly projected or barely projected forward, and distinctly projected forward in female; lateral angles absent; posterior angles barely visible or obtuse; margins of latero-basal third slightly convergent; lateral margin of anterior half very distinct, easily visible from above. Elytral punctation variable. Veins MP 3 and MP 4 not fused at their apex ( Fig. 210 ). Apex of prosternal process slightly enlarged. Femora glabrous or with hair very short and very sparse. Dorsal face of tibiae sulcate or slightly sulcate; outer face with longitudinal carina very distinct. Procoxal cavities distinctly open behind. Paronychium with one seta.

Included species. Papuandra araucariae ( Gressitt, 1959) , comb. nov.; P. gressitti Santos-Silva, Heffern and Matsuda , sp. nov.; P. weigeli Santos-Silva, Heffern and Matsuda , sp. nov.; P. queenslandensis Santos-Silva, Heffern and Matsuda , sp. nov.; P. norfolkensis Santos-Silva, Heffern and Matsuda , sp. nov.; P. rothschildi Santos-Silva, Heffern and Matsuda , sp. nov.; and P. oberthueri Santos-Silva, Heffern and Matsuda , sp. nov.

Geographical distribution ( Fig. 316 ). Indonesia (Irian Jaya), Papua New Guinea (New Guinea Island and Normamby Island), and Australia ( Norfolk Island).

Comments. Papuandra differs from Birandra by the mandibles sub-falciform or distinctly not falciform (distinctly falciform in Birandra ). Differs from Acutandra by the mandibles of the males not tumid at outer face, and by the dorsal face of the metatibiae longitudinally sulcate. In Acutandra , the mandibles of the male are tumid at outer face, and the dorsal face of the metatibiae is not sulcate. Differs from Archandra , Parandra , Stenandra , and Neandra by the procoxal cavities open behind (closed in the other four genera). Differs from Komiyandra by the margins of the latero-basal third of the prothorax slightly convergent, by the lateral margin of the anterior half of the pronotum very distinct and easily visible from above, by the veins MP 3 and MP 4 ( Fig. 210 ) not fused at their apices, by the dorsal face of the metatibiae longitudinally sulcate, and by the distinct presence of a carina on outer face of the metatibiae. In Komiyandra , the margins of the latero-basal third of the prothorax are clearly convergent, the lateral margin of the anterior half of the pronotum is not distinct, and frequently, is hardly visible from above (except in K. vivesi ), the MP 3 and MP 4 ( Fig. 211 ) are fused at their apices, the dorsal face of the metatibia is not longitudinally sulcate or is slightly sulcate only on apical half, and the lateral carina of the metatibia is visible or slightly visible. Differs from Melanesiandra by the outer margin of the mandible wide (narrow in Melanesiandra ). Differs from Hawaiiandra by the mandibles of the males not falciform and Birandra -like in females, and by the apex of prosternal process slightly wide at apex. In Hawaiiandra , the mandibles of the males are falciform, and Parandra -like in females, and the apex of prosternal process is wide apically. Differs from Caledonandra by the dorsal carina of the mandibles of the males not strongly oblique in relation to the longitudinal axis, and by the absence of depression in “V” at dorsal face of the head between the eyes. In Caledonandra the dorsal carinae of the mandibles of the male are strongly oblique in relation to the longitudinal axis, and there is depression in “V” at dorsal face of the head between the eyes. Differs from Malukandra by the absence of a tooth at the middle of the outer face of the mandibles (present in Malukandra ). Finally, differs from Storeyandra by the presence of a small tooth at apex of the outer margin of the mandibles in both sexes, and by the metathorax of the males not reduced (tooth absent and metathorax reduced in Storeyandra ).

Figures

Figure 75-89. Head, lateral view, male. 75) Hawaiiandra puncticeps. 76) Storeyandra frenchi. 77) Malukandra heterostyla. 78) M. jayawijayana, holotype. 79) M. hornabrooki, holotype. 80) Caledonandra austrocaledonica. 81) C. passandroides. 82) Melanesiandra striatifrons. 83) M. solomonensis, paratype. 84) M. bougainvillensis, holotype. 85) M. birai, holotype. 86) Papuandra araucariae. 87) P. gressitti, holotype. 88) P. weigeli, holotype. 89) P. norfolkensis, holotype.

Figure 385-390. Habitus. 385) Melanesiandra birai, holotype male, dorsal view. 386) M. birai, holotype male, ventral view. 387) Papuandra araucariae, male, dorsal view. 388) P. araucariae, male ventral view. 389) P. araucariae, female, dorsal view. 390) P. gressitti, holotype male, dorsal view.

Figure 118-147. Mandibles, dorsal view. 118) Birandra (Yvesandra) angulicollis, male. 119) B. (Y.) angulicollis, female. 120) Acutandra araucana, male. 121) Parandra (Parandra) glabra, female. 122) Stenandra kolbei, female. 123) Hawaiiandra puncticeps, male. 124) Storeyandra frenchi, male. 125) S. frenchi, female. 126) Malukandra heterostyla, holotype male. 127) M. jayawijayana, holotype male. 128) M. hornabrooki, holotype male. 129) M. hornabrooki, paratype female. 130) Caledonandra austrocaledonica, male. 131) C. austrocaledonica, female. 132) C. passandroides, male. 133) C. passandroides, female. 134) Melanesiandra striatifrons, male. 135) M. striatifrons, female. 136) M. solomonensis, male. 137) M. solomonensis, female. 138) M. bougainvillensis, holotype male. 139) M. bougainvillensis, paratype female. 140) M. birai, holotype male. 141) Papuandra araucariae, paratype male. 142) P. araucariae, female. 143) P. gressitti, holotype male. 144) P. weigeli, holotype male. 145) P. weigeli, paratype female. 146) P. queenslandensis, holotype male. 147) P. oberthueri, holotype female.

Figure 391-396. Habitus. 391) Papuandra gressitti, holotype male, ventral view. 392) P. weigeli, holotype male, dorsal view. 393) P. weigeli, holotype male, ventral view. 394) P. weigeli, paratype female, dorsal view. 395) P. queenslandensis, holotype female, dorsal view. 396) P. queenslandensis, holotype female, ventral view.

Figure 200-209. 200-204) Maxillary palp. 200) Komiyandra nayani, paratype male. 201) Papuandra araucariae, male. 202) Storeyandra frenchi, female. 203) Melanesiandra striatifrons, male. 204) Acutandra murrayi, female. 205-209) Wing. 205) Acutandra murrayi, female. 206) Neandra brunnea, male. 207) Archandra caspia, male. 208) Parandra (Parandra) ubirajarai, male. 209) Hawaiiandra puncticeps, male.

Figure 218-234. Antenna. 218) Malukandra jayawijayana, holotype male. 219) M. hornabrooki, paratype female. 220) Caledonandra austrocaledonica, male. 221) C. passandroides, male. 222) Melanesiandra striatifrons, male. 223) M. solomonensis, male. 224) M. bougainvillensis, holotype male. 225) M. birai, holotype male.226) Papuandra araucariae, male. 227) P. gressitti, holotype male. 228) P. weigeli, holotype male. 229) P. queenslandensis, holotype female. 230) P. norfolkensis, holotype male. 231) P. rothschildi, paratype male. 232) P. oberthueri, holotype female. 233) Komiyandra janus, female. 234) K. shibatai, male.

Figure 210-217. 210-214) Wings. 210) Papuandra araucariae, male. 211) Komiyandra nayani, paratype male. 212) Birandra (Birandra) silvaini, male. 213) Caledonandra passandroides, male. 214) Melanesiandra striatifrons, male. 215-217) Antenna. 215) Stenandra kolbei, female. 216) Hawaiiandra puncticeps, male. 217) Storeyandra frenchi, male.

Figure 315-316. Maps. 315) Distribution of Melanesiandra, Caledonandra, Storeyandra, Hawaiiandra, Malukandra, and Stenandra in Asia. 316) Distribution of Komiyandra and Papuandra.