Clivina schillhammeri, Balkenohl, 2021

Balkenohl, Michael, 2021, Belgian Journal of Entomology, Belgian Journal of Entomology 115, pp. 1-83 : 46-53

Clivina schillhammeri sp. nov.

urn:lsid:zoobank.org:act:D0FD6935-0961-4DBE-B9FB-49124FD3486F

( Figs 31 , 51 , 52 , 71 , 83 , 98 , 115 )

ETYMOLOGY. The species is dedicated to Dr. Harald Schillhammer (NHMW) who collected numerous specimens of the species in Laos and in Myanmar and who supported my work over many years.

TYPE MATERIAL. Holotype: • ♂, with labels and data: white, black printed “N- LAOS: Prov. leg. Nam Tha Muang Sing, at light 9–13.vi. 1996, 600 m leg. Schillhammer (18) ( NHMW) .

Paratypes: KOREA. • 1 ♀, KOREA: Prov. south Phenan: Pyongyang 4.vi.1970 / Hung. Zool. Exp. I. in Korea. No. 92., leg. Dr. S. Mahunka et Dr. H. Steinmann / Clivina castanea Westw. det. Alex. Dostal 1989 ( CADW) .

JAPAN. • 8 ♂♂, 8 ♀♀, Saeki, Oita Prev. 24.vii.1948 Kiushiu. Japan ( CADW) ; • 1 ♂, 1 ♀, same data but 5.viii.1948 ( CADW) ; • 2 ♂♂, 4 ♀♀, same data but 11.viii.1948 ( CADW) ; • 5 ♂♂, 12 ♀♀, same data but 28.viii.1948 ( CADW) ; • 1 ♂, Kobe, Hyogo. vii.1949 Honshin, Japan ( CADW) ; • 3 ♂♂, 1 ♀, Fujioka-chô, Tochigi Pref. / Wartarase-yusuichi / 8.ix.2007 T. Suda leg. ( CSMT) ; • 1 ♀, handwritten in black ink: Japon. Lewis / Clivina parryi Putz det. J. Putzeys / printed: Soc. Ent. Belge Coll. PUTZEYS ( RBINS) ; • 1 ♂, 1 ♀, handwritten in black ink and printed: Matsuyama Ehime Pref. 10.vi.1955 / R. I. Sc. N. B. I. G. 22.024 ( RBINS) ; • 1 spec., same data but 30.vi.1954 ( RBINS) .

TAIWAN. • 1 ♂, 1 ♀, TAIWAN, Pingtung County Kenting Liu-fu Hotel N21°57.388 E 120°47.017, 02.viii.2008, at light H. Mendel, MVL Barclay & R. Ewers BMNH(E)2008–85 ( BMNH) ; • 1 spec., TAIWAN, Lanyu Island (Taitung County) Yung Hsin Farm N 22°01.703 E 121°34.642, 29–30.vii.2008, at light H. Mendel & MVL Barclay BMNH(E)2008-85 ( BMNH) ; • 1 ♂, S. Formosa Kagi 20–viii.1907 H. Sauter S.V. No. 11474 ( MFNB) ; • 1 spec., TAIWAN vii.1990 Pingtung Hsien Suchung-Shi / 44 leg. M.L. Jang ( NHMW) ; • 1 spec., Kosempo Formosa H. Sauter 22.v.1912 / Clivina castanea West. det. Ing. Jedli Č ka / ex Coll. Jedli Č ka Nat. Mus. Prague ( NHMP) .

HONG KONG. • 1 ♂, Hong Kong, China coll: Paul Aston 2–9.x.1912 Pui O, Lantau. H.K. PT in Marsh / Clivina castanea Westwood 1837 Determined D. Fedorenko ( CMBB) ; • 2 ♀♀, HONGKONG, 1983 Tai Po Kau, 10.viii. downstream leg. Dudgeon ( NHMW) .

CHINA. • 1 ♂, Kiautschau China / C. castanea / COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 ( CADW) ; • 1 spec., Kiautschau China /? moerens Putz. Det. Fleischer ( NHMP) ; • 1 ♀, China, Chinkiang 32°12’N 119°30’E, 4.i.1919 Eigin Suenson leg. ( CADW) ; • 2 ♂♂, China, Soochow 31°21’N 120°40’E, 6.iv.1919 Eigin Suenson leg. ( CADW) ; • 2 ♀♀, China, Shanghai 31°13’N 121°25’E 6.v.1937 Eigin Suenson leg. ( CADW) ; • 1 spec., Zi-ka-wei 16.vi.20 / O. PIEL coll. ( CADW) ; • 1 ♂, 1 ♀, China, Hangchow 30°18’N 120°07’E, 18.x.1920, Eigin Suenson leg. ( CADW) ; • 1 ♂, China, Canton, Pak wan Asi , 15.vi.1910 Mell S. V. ( MFNB) ; • 1 spec., China Canton /? moerens Putz. Det. Fleischer ( NHMP) ; • 1 spec., Tsangen /? moerens Putz. Det. Fleischer ( NHMP) ; • 1 spec., Shaowu – Fukien (500m) J. Klapperich / Clivina castanea West. Dr. Maran det. ( NHMP) ; • 3 spec., KUATUN FUKIEN China meridion. Klapperich / castanea Westw. det. Ing. Jedli Č ka / ex Coll. Jedli Č ka Nat. Mus. Prague ( NHMP) ; • 1 ♀, Südchina Wuhan 15.viii.1984 Katschok leg. / Clivina castanea Westw. det. Balkenohl 1999 ( SMNS) ; • 1 ♂, CHINA – Guangdong Chebaling Nat. Reserve E. Shaoguan , 300–500 m 25–27.vi.1996 Chr. Häuser leg ( SMNS) ; • 6 ♂♂, 3 ♀♀, China, Buong Kä n. of. Foochow – 27°N 119°E, 26.iv.1935 Eigin Suenson leg. ( CADW) ; • 1 ♂, China, Kushan 26°4’N 119°25’E 8.iv.1935 Eigin Suenson leg. ( CADW) ; • 1 spec., KATUN , FUKIEN China 4.iv.1946, (TSCHUNG SEN.) ( CADW) ; • 2 ♂♂, 1 ♀, China, Kuliang 26°12’N 119°28’E 11.iv.1935 Eigin Suenson leg. ( CADW) ; • 1 ♀, SO China: Fujian Wuyi Gebirge (ca 125 km N Yvanping 200– 400 m 7–22.iv.1988 leg. Görgner & Kleinfeld ( CWGP) ; • 1 ♀, China, Nanping 26°50’N 118°05’E 17.v.1935 Eigin Suenson leg. ( CADW) ; • 1 spec., Kiang-Si T‘ en- gan ( MFNB) ; • 1 spec., same data and ex Coll. Jedli Č ka Nat. Mus. Prague ( NHMP) ; • 1 ♂, CHINA, N GUANGXI reg. MIAOERSHAN , S slope, 25–26.vi.1997, 1300–2000 m, Bolm lgt / CLIVINA (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NHMB) ; • 1 spec., Chine centrale SI ‘ Shan 25.v.1931 R.P. Marga / R. Mus. Hist. Nat. Belg.I.G.13.336 / Clivina sp. det. S.W.Nichols 1984 / Clivina castanea s.l. Westwood, 1837 P. Bulirsch det. 2011 ( RBINS) ; • 1 spec., CHINA, ZHEJIANG Prov., West Tianmu Shan (Mts) reserve, border of secondary mixed forest nr entrance of reserve; at light 30°18´34.5“N, 119°28´46“E, 310 m, 23.vi.–6.vii.2017 J. Hájek & J., Růzička leg. ( NHMP) ; • 1 ♀, Kweichou China, Plason ( CADW), • 1 ♂, CHINA / Sichuan 103.20el/29.30 NW Mt. Emei 500–1200 m 4–19.v.1989 S. & J. Kolibéč leg. / Freiwilliger Museumsverein Basel 1989 ( NHMB) ; • 2 ♂♂, 1 ♀, 10 specs., CHINA (Yunnan) Lincang Pref., Mekong Valley , small creek cleft 38 km SSE Lincang . 854 m 23°33’13.2”N / 100°09’56.8”E (field edge in small cleft, in loamy soil) 11.ix.2009 D.W. Wrase [44] / Coll. Wrase Berlin ( CWGP, CMBB) ; • 1 ♀, same data but ( NKME) ; • 1 spec., China, Prov. Yunnan Gbg. B. Mengtze / Clivina castanea Westw. H.E. Andrewes det. / ex Coll. Jedli Č ka Nat. Mus. Prague ( NHMP) ; • 1 spec., CHINA: YUNNAN Prov. Tongbiguan vill. 24–26.vi.2016 24°36.7’N, 97°39.4’E, 1340 m at light in village near river J. Hájek & J., Růzička leg. / CLIVINA castanea Westwood, 1837 , P. Bulirsch det. 2018 ( NHMP) ; • 1 spec., Tsushima Rost 1919 / ex Coll. Jedli Č ka Nat. Mus. Prague ( NHMP) ; • 3 historic specs. with black framed and printed labels, one without abdomen: CHINA ( NHMB) ; • 1 historic spec. with double framed and handwritten label: Lilong. China ( NHMB) ; • 1 ♂, Vallis flumin. Soling-ho, Yun . / castanea Westw. det. Ing. Jedlička / A 1975 / WIEN/COLLECTIO Dr. R. WEBER; 1994 ( CADW) ; • 1 spec., Tchenkiang / Lou bou 10.vii.1923 / O. PIEL, coll. ( CADW) ; • 1 spec., Kiang-si Kiu-kiang ( MFNB) ; • 1 spec., Kiang-si Chang-tsin-cheng ( MFNB) ; • 1 spec., Tientsin 16, IX. 29 / castanea Westw. det. Ing. Jedlička ( NHMP) .

VIETNAM. NORTH VIETNAM. • 1 ♂, Pa-kha 1914 / Indo Chine Coll. Dussault ( NHMB) ; • 1 spec., VIETNAM SaPa – 1530 m 25.v–9.vi.1991 Strnad Jan leg. ( CMBB) ; • 1 spec., N-VIETNAM 1991 75 km NW Hanoi Tam Dao 16–23. v. leg. J. Strnad ( NHMW) ; • 1 ♀, Annam Phuc Son ( RBINS) ; • 2 specs., Annam Phuc son Nov. Dez. H. Fruhstorfer ( MFNB) ; • 2 ♂♂, Annam Phuc-Son Nov. Dez. H. Fruhstorfer ( MFNB) ; • 1 spec., N-Vietnam Vinh Phu pr. Tam Dao 1050–1200 m 21°27.18N 105°38.58E 2–6.vi.1999, leg. Ahrens ( CJSR) ; • 3 specs., Coll. I.R.Sc.N.B. VIETNAM: Phia Den 08.vii ..2010, Light Trap I.G. 31.668 leg. J. Constant & P. Limbourg ( RBINS) ; • 1 spec. Hanoi 1923/25 / Indo Chine Coll. Dussault / Clivina castanea WESTW. det. Balkenohl xi.1991 ( NHMB) ; • 1 ♂, 10 specs., N-Vietnam Lao Chai pr. Bac Ha env. 980–1000 m 22°22,05N 104°17.32E 28–30.v.1999, leg. Ahrens ( CJSR) ; • 1 ♀, Tonkin Hanoi / castanea Westw. det. Ing. Jedlička / castanea Hoppe [sic] det. K. Kult / COLLECTIO KAREL KULT COLL. A. DOSTAL, 1999 ( CADW) ; • 2 ♂♂, TONKIN Hoa-Binh , vii.1940, leg. A. de Cooman ( CADW) ; • 1 spec., same data but iii.1939 ( CADW) ; • 2 specs., same data but iii.1931 ( CADW) ; • 1 ♂, TONKIN Hoa-Binh leg A. de Cooman / No 3 ( CADW) ; • 1 spec., Hoah Binh Tonkin / Clivina castanea WESTW. det. M. Baehr’ 05 ( CMBM) ; • 1 spec., Vietnam, Hoa binh 4–7.vi.1986 Ha son binh prov. Jan Horák leg. / CLIVINA castanea group P. Bulirsch det. 2009 ( NHMP) ; • 1 spec., VIETNAM-N, Tuyen Quang pr. SE-E env. of Na Hang 22°17’30”- 22°30’N 105°26’ –28’E, 200–700 m L. Dembický leg., 1–12.v.2010 / CLIVINA (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NHMB) ; • 1 ♂, N VIETNAM (Tonkin) pr. Hoang Lien Son SAPA 11–15.v.1990 Vít. Kubán leg. ( CADW) ; • 1 ♀, Kansou mer. Hoei-sien. ( MFNB) ; • 2 specs., VIETNAM N, Cuc Phuong 2–11.v.1991 Strand Jan leg. ( NHMB) ; • 1 spec., N-VIETNAM Ninh Binh Cuc Phuong NP, 270 m N 20°17’57’’, E 105°40’05’’, 04–09.v.2017. A. Weigel KL / collection NATURKUNDE-MUSEUM ERFURT / CLIVINA (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2018 ( NKME) ; • 1 spec, same data but 25.iv. 2012, 190 m, N20°14’24’’, E105°42’53’’ leg. A. Weigel Li.-Fa. / CLIVINA (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2014 ( NKME) ; • 2 specs., N-VIETNAM Cao Bang Pr. vic. Vin Den, Nui Pia Oac Nature Res. , 6–10.v.2013, 22°33’53’’N, 105°52’53’’E 900–1300 m leg. A. Weigel / collection NATURKUNDE-MUSEUM ERFURT / CLIVINA (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2014 ( NKME) ; • 1 spec., N-VIETNAM Thai Nguyen Pr. vic. Ngoc Thanh, vic. Me Linh ( IEBR station ), 12.v.2012, 21°22‘3’’N, 105°42’44’’E 60–80 m leg. A. Weigel / collection NATURKUNDE-MUSEUM ERFURT / CLIVINA (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2014 ( NKME) ; • 3 specs., VIETNAM, N (Na Hang) 160 km NNW Hanoi NE env. of Na Hang 28.v.–10.vi.1996, LF, 150–200 m, NN, leg. A. Napolov & I. Roma / collection NATURKUNDE-MUSEUM ERFURT / CLIVINA (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2012 ( NKME) ; • 1 ♀, (305) Annam-Vinh 1910 / Indo Chine Coll. Dussault ( NHMB) ; • 1 spec., VIETNAM: Me linh Light Trap 20–24.viii.2010, leg J. Constant & P. Limbourg I.G.31.668 ( RBINS) ; • 1 ♀, same data as before ( CMBM) .

SOUTH VIETNAM. • 1 ♂ [immature], VIETNAM: Buon-loi 40 km Nof Ankhe Prof. Gialai Kontum 15.vi.1985 ( SMNS) ; • 2 specs., S-VIETNAM 40km NW An Khe Buon Luoi, 620– 750 m / 14°10’N 108°30’E 28.iii.–12.iv.1995 Pacholatko & Dembicky ( NHMW) ; • 1 ♂, 4 specs., S-VIETNAM Nam Cat Tien Nat. Park 1–15.v.1994 Pacholatko & Dembicky ( NHMW) ; • 1 spec., VIETNAM Buon-loi 40 km Nof Ankhe Prov. Gia-lai Kontum 15.vi.1985 ( CMBB) .

THAILAND. • 1 ♀, THAILAND bor. CHIANG DAO env. 17–19.vi.1995 leg. SNIZEK M. ( CMBB) ; • 2 ♀♀, NO-THAILAND Nan – Pha Khab 11-16.v.1993 leg. Pacholatko ( NHMW) ; • 5 ♀♀, NO – THAILAND 1993 NAN – PHA 11-16. v. leg PACHOLATKO ( CADW) ; • 1 ♂, O – THAILAND 1990 Ko Chang (11) am Licht leg. Jäch 8– 12.xii. ( NHMW) ; • 1 spec. THAIL. 9.xii.1990 Ko Chang NP, leg. Forster ( NHMW) ; • 1 spec., E Thailand, 5–13.v.1998, Chanthaburi Dist. Khao Soi Dao , J. Fyman lgt ( NHMP) ; • 1 spec. Nord-Thailand Chieng Mai Dr. Jaratpun Sa-nguansermsri leg ix.1981 ( MFNB) ; • 1 ♂, NW THAILAND, Pai, 500 m, 17– 19.iv.2004, leg. W. SCHAWALLER ( SMNS) ; • 1 ♀, NW-THAILAND, CW Mae Hong Son Pai Thapai Hot Springs 25.iv.2004, R. Grimm ( SMNS) ; • 1 ♂, THAILAND PAI env. 1– 14.v.2001, J. Hromádka leg. ( CADW) ; • 1 ♂, 1 ♀, NW Thailand MAE HONG SON 1992 Nupa-ah 7– 9.v, Strnad Jan leg. ( CADW) ; • 1 ♂, THAILAND: Changwat Mae Hong Son. Ban Huai Kha 5.i. 1989, 300 m leg. TRAUTNER & GEIGENMÜLLER ( SMNS) ; • 1 spec., Thai, 26–28.v.1991, PALONG 750 m 19°55’N 99°06’E Vít Kubáň leg. / Thailand ’91 “Thanon Thong Chai” D. Král & V. Kubáň ( NHMB) ; • 1 spec., Coll. I.R.Sc.N.B. THAILAND (loei) Na-Haeo (field res stat) 15–19.v.2003 Light trap, leg. J. Constant, K. Smets & P. Grootaert ( RBINS) ; • 2 specs., THAI, 9–13.iv.1991, THIMONGHTA 350 m, 15°02’N 98°35’E Vít Kubáň leg. / Thailand ’91 “Thanon Thong Chai” D. Král & V. Kubáň ( NHMB) ; • 1 spec., Thai 18–24.iv.1991 LANSANG nat. park 16°48’N 98°57’E leg. D. Král 500 m / Thailand’91 “Thanon Thong Chai” D. Král & V. Kubáň / Clivina castanea WESTW. det. Balkenohl, 2007 ( NHMB) ; • 1 ♂, THAI 20.v.1993, NAKHON THAI Pacholatko & Dembicky leg. ( CADW) ; • 1 ♂, 1 ♀, 1 spec., THAILAND, S 40 km NE OF BETONG, LAKE BANG LANG (KG.TO) 12–23.vi. 2007, 350 m Z. LINEK leg. (CADW) ; • 1 spec., S-THAILAND Taku Pa , 20.xi.1996, leg. J. REJSEK ( SMNS) .

LAOS. • 1 ♀, same data as holotype ( NHMW) ; • 12 specs., LAOS, Phongsaly prov., PHONGSALY env., 6–17.v.2004,~ 1500 m, 21°41’N 102°06–08’E, M. Brancucci leg. / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NHMB) ; • 1 ♀, LAOS, PHONGSALI PROV. MEUNG KHOA DISTR. BAN HOUAYWANG 10–17.vi.2010 leg. SASA KAMPHILAVONG ( CADW) ; • 1 ♀, LAOS-Houaphan prov. 38 km Sof Sam Neua Saleui 9–22.v.2009 Martinu leg. 1350–1900 m ( CADW) ; • 3 ♀♀, NE-LAOS Hua Phan prov. Ban Saleui, Phou Pan Mt. 20°12’N 104°01’E 23.iv.–15.v.2008, 1500–1900 m, leg. C. Holzschuh ( CADW) ; • 1 ♀, same data but 11.iv.–15.v.2012, 1300–1900 m / collection NATURKUNDE-MUSEUM ERFURT / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2013 ( NKME) ; • 1 ♂, 6 specs., same data but 01–31.v.2011, leg. C. Holzschuh and local collector ( NKME) ; • 4 specs., LAOS: N-VIENTIANE Prov. Vang-Vieng, 300 m N 18°55’23”, E 102°26’55” 10–15.v. & 01–06.vi.2001, JIŘI KOLIBÁČ leg. ( NHMB) ; LAOS, Vientiane prov. LAO PAKO env. 200 m 55 km NE Vientiane 19–22.v.2004, J. Bezdĕk leg. / CLIVINA castanea group P.Bulirsch det. 2009 [sic] ( NHMP) ; • 1 ♂, 1 ♀, LAOS, Xieng Khouang Prov., 19°37-8’N 103°20’E, Phonsavan (30 km NE); Phou Sane Mt. 1400–1700 m 10–30.v.2009, D. Hauck leg. ( NHMB) ; • 4 specs., LAOS, Khammouan prov. 17°57 -59’N/ 105°13 -16’E, Nakai- Nam Hheun NPA, BAN NAVANG env., 600–750 m, 18–21.v.2012, / NHMB Basel, Expedition Laos 2012: M. Brancucci, M. Geiser, K. Phanthavong, S. Xayalath / Clivina (Leucocara) castanea Westwood, 1837 , P. Bulirsch det. 2016 ( NHMB) ; • 3 specs., LAOS, Bokeo prov., 5 km W Ban Toup, Bokeo Nature reserve , 500–700 m, 20°27 –28’ N/ 100°45’ E, 4–18.v.2011 / NHMB Basel, Laos 2011 Expedition, M. Brancucci, M. Geiser, D. Hauk, Z. Kraus, A. Phantala, & E. Vongphachan / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NHMB) ; • 2 specs., LAOS Nam Tha le 9.iv.1918, R. Vitalis de Salvaza / Clivina castanea West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 ( RBINS) ; • 6 specs., same data but Pak Pha 4.ii.1918 ( RBINS) ; • 1 spec., LAOS Pak Pha 4.ii.1918, R.V.de Salvaza / Clivina castanea Westw., H.E. Andrewes det. / Ex Misc Coleoptera 142 Tribes of the Carabidae Hope Ent. Coll. Oxford ( OUMNH) ; • 1 spec., LAOS Ehalan le 20.xi.1917, R. Vitalis de Salvaza / Clivina castanea West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 ( RBINS) ; • 1 spec., LAOS Ban Hou le 7.x.1918, R. Vitalis de Salvaza / Clivina castanea West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 ( RBINS) ; • 1 spec., LAOS Muong Pek le 15.xii.1918, R. Vitalis de Salvaza / Clivina castanea West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 ( RBINS) ; • 1 spec., LAOS B. Honei Soni le 30.i.1918, R. Vitalis de Salvaza / Clivina castanea West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 ( RBINS) ; • 1 spec., LAOS Natung le 29.iii.1918, R. Vitalis de Salvaza / Clivina castanea West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 ( RBINS) ; • 4 specs., LAOS-C: KHAMMOUAN pr., Ban Khoun Ngeun. ~ 200 m; 18°07’N 104°29’E; Pacholátoko leg., 24–29.iv.2001 / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch , det. 2016 ( NHMB) ; • 2 specs., LAOS, Salavan prov., 15°42’N / 106°24’E, SALAVAN town, 200 m, 23–24.v.2012, M. Geiser leg. / Collection Naturhistorisches Museum Basel / Clivina (Leucocara) castanea Westwood, 1837 , P. Bulirsch det. 2016 ( NHMB) ; • 2 specs., LAOS centr., Kammouan prov. NAKAI env. 4–8.v.1998, Route No 8 , alt. 560+ 20 m N 17°42.8, E 105°08.9 (GPS), M. Strba & R. Hergovits ( CWGP) ; • 3 specs., LAOS, Salavan prov., 16°08’N / 106°40 -43’E, Xe Xap NPA, ca. 15 km NE of Ta-oy , BAN DOUB env. 400–1000 m, 25–31.v.2012, / NHMB Basel, Expedition Laos 2012: M. Brancucci, M. Geiser, K. Phanthavong, S. Xayalath / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NHMB) ; • 3 specs., LAOS-N (Oudonxai), 1–9.v.2002, ~ 1100 m, 20°45’N 102°09’E, OUDOM XAI (17 km NEE). Vít Kubáň leg. / Collection Naturhistorisches Museum Basel / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NHMB) ; • 1 spec., LAOS-N(Louangphra-bang), 1–21.v.2002, 19°35’N 101°58’E, THONG KHAN, ~ 750 m, Vít Kubáň leg. / Collection Naturhistorisches Museum Basel / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NHMB) ; • 1 spec., LAOS, Champasak Prov., 15°02’N 106°31 -34’E, Ban Nong Panouan env. 770–800 m 10– 17.vi.2010, M. Geiser & D. Hauk leg. / coffee plantations, secondary and degrades primary forest; hand collecting, sweeping, beating, coll. at flowering trees and in small water puddles, light trapping ( NHMB) ; • 1 spec., LAO, 26–27.v.2003, Bolikhamxai prov., PAKKADING, ~ 300 m, 18°20’N 104°00’E, Vít Kubáň leg. / Collection Naturhistorisches Museum Basel / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2016 ( NHMB) ; • 3 specs., S-LAOS: Prov. Champasak Muang Paksong, at light 26–27.v.1996, ca. 1100 m, leg. Schillhammer (8) ( NHMW) ; • 1 ♀, S-LAOS, Pr. Champasak, Bolavens-Plateau (E Pakse) Muang Paksong – Ban Nam Thang , 1000 m, 26–28.v.1996, leg. C. Holzschuh. / collection NATURKUNDE-MUSEUM ERFURT / Clivina (Leucocara) castanea Westwood, 1837 P. Bulirsch det. 2017 ( NKME) . • 3 specs., LAOS n., 13–24.v. 1997, 15 km NW Louang Mantha , alt. 750–1000 m N21°07.5’; E101°21.0 ǴSP, E. Jendek & O. Sausa leg. ( NHMP) ; • 2 specs., LAOS, SEKONG prov. ca. 12 km S Sekong TAD FAEK waterfalls (at light) 15°14.7’N, 196°45.1’E, 188 Jiří Hájek leg. 8–12.v.2010 / Clivina castanea s.l. Westwood 1837 P. Bulirsch det. 2010 ( NHMP) .

MYANMAR. • 1 ♂, MYANMAR, KACHIN PROVINCE, MT EMAW BUM 5–22.vi.2013 LOCAL COLLECTOR / trachys Andr. DET. DR. A. DOSTAL 2015 ( CADW) ; • 1 ♀, BIRMANIA Rangoon vi.52 Bentoglio ( CMBB) ; • 4 specs., Bhamo Birmania Fea iv.1886 ( RBINS) ; • 1 ♂, 1 ♀, 3 specs., MYANMAR: Sagaing Division Chatthin Wildlife Sanctuary 23°33.017’N 95°34.851’E Kinsan Camp , 210 m, 13.x.1998 leg. Schillhammer (14) ( NHMW) ; • 1 spec., MYANMAR: Shan State road Kalaw – Thazi, km 26 20°43.796’N 96°29.477’E 550 m, 24.x.1998, leg. Schillhammer (28) ( NHMW) ; • 1 ♂, BURMA ( MYANMAR) PEGU CHANTHAKWIN KLÍCHA M. leg. / Reichardtula castanea Westw. det. Dr. A. Dostal 2007 ( CADW) ; • 1 spec., MYANMAR (Kachin State) road Bhamo to Shwegu Irrawaddy river 157 m (light) 24°07’387”N / 97°01’577”E 5.vi.2006, M. Langer, S. Naumann & S. Löffler / Clivina castanea Westwood, 1837 P. Bulirsch det. 2005 / COLL. WRASE BERLIN ( CWGP) . • 1 ♀ [right hind leg missing] with labels and data: pale beige, black printed and handwritten in black ink “Coll. Helfer. Ind. or” / handwritten in black ink “ Clivina Parryi Putzeys Eupalamus clivinoides Schmidt-Goebel. ” / black printed “TYP. Schmidt-Göbel.” / black printed and handwritten in black ink, black framed “Mus. R. Boh. Inv. č. Clivina Parryi clivinoides S.G ” ( NHMP) ; • 1 ♀ [right hind leg dislocated], with labels and data: pale beige, black printed and handwritten in black ink “Coll. Helfer. Ind. or” / handwritten in black ink “ Clivina Parryi Putzeys Birmanie” / black printed “TYP. Schmidt- Göbel.” / black printed and handwritten in black ink, black framed “Mus. R. Boh. Inv. č. Clivina Parryi clivinoides S.G”. ( NHMP) . • 4 specs, MUS.PRAGENSE TENASSERIM Coll. HELFER ( NHMP) .

CAMBODIA. • 1 ♀, CAMBODGE / Clivina castanea Westw. 30.4.1930 H.E. Andrewes det. ( ETHZ) ; • 1 ♀, CAMBODIA SW 20 km SE KOH KONG, TATAI RIVER , 11°34’N 103°07’E, 17.x.–15.xi.2011 50–300 m, Z. LINEK LGT. ( CADW) ; • 1 spec., CAMBODGE Komp. Kedey le Mars 1914, R. Vitalis de Salvaza / Clivina castanea West. det. R. Vitalis de Salvaza / R.I.Sc.N.B 16.493 ( RBINS) ; • 1 spec., Coll. I.R.Sc.N.B. Cambodia – 8 km north of Sre Noi ( road to Along Vaeng ) Light trap, 29.v.2003, J. Constant & K. Smets ( RBINS) .

MALAYSIA. • 4 ♂♂, 6 ♀♀, MALAYSIA W, Pahang 70 km SW of KUALA ROMPIN , ENDAU ROMPIN NP 699 m G. BEREM-BAN (KG.TEBU HITAM) 13.iv.–03.v.2009 P.CECHOVSKY leg. ( CADW) .

INDIA. • 1 ♀, INDIA, 29.vi.–02.vii.1995 Umran 40 km S Shillong Meghalaya WERNER leg. ( CMBB); • 1 ♂, assam Lesid 94 / Clivina Parryi Putz. M. Maindron det. 1910 / Collection E. Rousseau ( RBINS) .

Paratypes of Clivina khasei Jedlička : • 1 ♂, with the following labels and data: white, printed “ ASSAM, Khasi- Hills , Cherrapunji ca. 1300 m, v.1961 leg. G. Scherer ” / “white, black printed “Museum Frey Tutzing” / red, black printed “Paratypus” / red, black printed and handwritten with black ink “ Clivina khasei sp. n. det. ING. JEDLIČKA ” / white, black printed “COLLECTIO KAREL KULT COLL.A.DOSTAL,1999” ( CADW) ; • 1 ♀, with the following labels and data: white, printed “ ASSAM, Kaziranga nördl. Mikir- Hills Brahmaputra , v.1961, leg. G. Scherer ” / “white, black printed “Museum Frey Tutzing” / red, black printed “ Paratypus ” / red, black printed and handwritten with black ink “ Clivina khasei sp. n. det. ING. JEDLIČKA ” ( NHMB) .

[Note: The specimen labelled as holotype with labels Clivina khasei Jedlička is listed among the faunistic data of Clivina lata Putzeys. ]

DIAGNOSIS. Amedium sized species with smooth lateral margin of the pronotum and elytron, and isodiametric reticulation on interval eight of the elytron. Distinguished from the other species with isodiametric reticulation on interval eight by the mentum with semi-oval shape of lobe and the complete and wide epilobe which are separated from the lobe by a carina. The only other species in which the lobes of the mentum share similarities is C. hogani sp. nov., but in that species interval eight is smooth. Moreover, the aedeagus is different.

DESCRIPTION. Measurements. Holotype: Body length 8.41 mm, width 2.47 mm; ratio length/width of pronotum 0.95; ratio length/width of elytra 1.89. Other material: Body length 7.95–9.02 mm (: = 8.51 mm *), width 2.26–2.63 mm (: = 2.47 mm *), ratiolength/width of pronotum 0.91–0.95 (: = 0.93*), ratiolength/widthofelytra 1.89–1.97 (: = 1.93*); (*n = 10).

Colour: Glossy. Piceous. Antenna and tarsomeres fuscous.

Head: Athird narrower than pronotum. Clypeus slightly excised anteriorly; wing slightly less protruding as clypeus, flattened, separated from clypeus by distinct notch; supraantennal plate convex; clypeus, and supraantennal plate reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch. Clypeus slightly transverse, slightly convex, separated from frons by moderately deep furrow. Frons moderately convex, with flat pore at middle, with moderately sized carina at posterior eye-level. Clypeus and frons smooth, with minutely scattered punctures, separated from wing and supraantennal plate by moderately deep and wide furrow continuing posteriorly as wide supraorbital furrow with two supraorbital setigerous punctures at posterior eye level. Neck constriction developed as distinct step, with row of small punctures. Eye semi-hemispherical projecting. Gena short, indistinct. Antenna reaching distinctly over posterior angle of pronotum, antennomeres four to ten elongate (ratio L/W around 1.95). Labrum nearly straight, with indistinct obtuse angle at middle, with isodiametric reticulation, seven setose. Mentum ( Figs 51 , 52 ) with distinct isodiametric reticulation, with lobe slightly hollowed out, shape of lobe semi-oval, epilobe wide and complete, separated from lobe by carina, with distinct carina at middle, median tooth wide, slightly less protruding anteriorly as epilobe, form like an anomalous pentagon.

Pronotum ( Fig. 71 ): Disk distinctly convex in lateral and frontal view. Slightly wider than long. Reflexed lateral margin smooth, regularly convex, narrowed anteriorly, widest in posterior third; anterior angle obtuse; posterior angle distinctly marked but laterally not as wide projecting as lateral margin in posterior third. Lateral channel stepped from disk at level of anterior setigerous puncture, with indistinct reticulation. Median line moderately wide, linelike, just joining basal constriction; anterior transverse line punctured, joining median line. Surface smooth, with some longitudinal and transverse wrinkles, with indistinct microscopic punctures, with isodiametric reticulation at base and laterally in basal half.

Elytron: Slightly depressed on disk in anterior half in lateral view, convex in frontal view. Nearly twice as long as wide; lateral margin slightly convex but not dilated, maximum width at middle. Reflexed lateral margin smooth. Scutellar striole distinct, slightly carinate; setigerous tubercle at base of first stria, with small but distinct tubercle at base of third interval. Striae moderately deep, punctuate-striate, one to three free at base, four to seven joining at humerus, one joining indistinctly with seven at apex, two to six ending free at apex, often stria five and six joining apically. Intervals moderately convex, fourth slightly wider at base, eights with short carina at humerus and longer one apically. Third interval with four setigerous punctures, first situated at second stria near base, the other three approaching third stria. Surface of interval eight finely isodiametric reticulated (80 x), the other intervals glossy, with reticulation at apical tip.

Hind wings: Fully developed.

Lower surface: Proepisternum covered by about half with isodiametric reticulation, with few fine transverse wrinkles ( Fig. 83 ). Terminal sternite of abdomen developed apically as flat narrow transverse fold.

Legs: Legs covered with indistinct reticulation. Protibia with three spines of moderate length, sulcate dorsally, movable spur slightly more arcuate at tip.

Male genitalia ( Fig. 98 ): Median lobe long, slender, narrowed at middle, broadened apically, slightly sinuate before apex, regularly tapered to and at apex, ventral rim distinct. Endophallus laterally with longitudinal group of around 18 spines. Ventral paramere distinctly widened, sinuate, lateral apophysis long, apex spatula-like tapered, with four long setae at apex. Dorsal paramere slightly and regularly arcuate, slender, at apex with three long setae and one short seta preapically.

Female coxostylus and laterotergite ( Fig. 115 ): Coxostylus moderately broadened at base, gently curved to apex (lateral view), slightly widened in apical half (ventral view), dorsally with eleven long and slender and one elongated and robust nematiform setae in basal two thirds, ventrally with numerous smaller setae, with two SSOs. Laterotergite with five long and one small setae basally and one isolated seta towards coxostylus.

VARIATION. The semi-oval lobe of the mentum is slightly narrower and a bit shorter in specimens from Japan ( Fig. 52 ) but becoming continuously and slightly wider in specimens from Eastern to Western China. The reticulation at the apical tip of the elytron is slightly varying. In few specimens striae five and six joining at apex at one side. The flat central pore on the frons of the head is missing in a few specimens, in others it is distinct. In two males the dorsal paramere shows an additional very small seta at the apex. The proepisternum is in most of the specimens covered by about half with isodiametric reticulation but in some specimens up to 75 %. Variation was also observed in the isodiametric reticulation of interval eight of the elytron which covers half of the width of the interval in some specimens and is extended to the whole interval in other specimens. In specimens from China and Vietnam it is often weak and requires high resolution (min. 100x) to be visible. Specimens from the North of Thailand, Laos, Myanmar and the middle of Vietnam do not show reticulation on interval eight at all (160×).

DISTRIBUTION: Korea, Japan, Taiwan, Hong Kong, China, North and in the middle of Vietnam, North Thailand, Myanmar, Cambodia, and up to the North of Malaysia.

Figures

Fig. 31. C. schillhammeri sp. nov., holotype.

Fig. 51. C. schillhammeri sp. nov., holotype.

Fig. 52. C. schillhammeri sp. nov., paratype from Japan (Waterase-yusuichi Fujiokachô).

Fig. 71. C. schillhammeri sp. nov., holotype.

Fig. 83. C. schillhammeri sp. nov., holotype.

Fig. 98. C. schillhammeri sp. nov., holotype.

Fig. 115. C. schillhammeri sp. nov., paratype.